EEPORT ON THE CRINOIDEA. 215 



With regard to the support beneath the radial pentagon of Eudesicrinus, I see no, 

 reason for doubting that the upper part consists of united basals. In fact, one of de 

 Loriol's figures* shows a distinct horizontal suturalline crossing the middle of the support, 

 and separating the infra-radial portion with interradial crests on its upper surface from 

 the more spreading, attached part below. In another specimen this suture seems to be 

 indicated by an external circular ridge ; but the upper face of the support is marked by- 

 five petaloid depressions, one of them considerably larger than the rest, which surround 

 a central pit. De Loriol, and, I think, rightly so, regards these depressions as corresponding 

 to the cavities of the chambered organ, the largest being that of the large radial in the 

 trivium. Similar but more regular depressions appear round the middle of the upper 

 surface of the large basal pentagon in Apiocrinus milleri, Quenstedt ; and they are also 

 shown in de Loriol's figure af Ajnocrinus roissyanus,^ while it will be remembered that 

 the chambered organ is invariably in close relation with the basals (PL Vllb. figs. 1,2; 

 PL XXIV. figs. 6, 7 ; PL LVIII. figs. 1, 3—ch; PL LXL). There can then, I think, be 

 no doubt as to the presence of basals in Eud-esicrinus, so that the so-called support does 

 not in reality difi"er essentially from the centro-dorsal of Cotylecrinus. The radials, how- 

 ever, are very different in the two cases. Those of Cotylecrinus are equal and similar ; 

 but in Eudesicrinus they are thus described by de Loriol,' " Ces pieces sont fort inegales ; 

 I'une est notablement plus longue et plus large que les autres, convexe et un pen arqude 

 en dehors, mais d'une maniere uniforme ; les deux qui la touchent, bien plus courtes et plus 

 etroites qu'elle-meme, ne sont guere plus larges, mais plus longues que les deux autres ; 

 ces dernieres, qui sont les plus courtes et placees vis-k-vis de la plus large, s'arquent en dedans 

 et se coudent un peu vers leur bord superieur. Dans les echantillons frais la surface externe 

 est couverte de granules epars, ecartes, plus ou moins gros et plus ou moins serrds." 



Here then we have a type which bears an unusually close resemblance to Holoims. 

 The calyx is higher on oiie side than on the other o-ndng to the inequality of the radials, 

 the central one of the trivium being the largest, while the outer surface, not only of the 

 radials, but also of the arm-j oints, is coarsely granular or tul^ercular. Eudesicrinus, however, 

 difi"ers from Holopus in one or two minor points. The fossse on the radials which lodged 

 the dorsal ligaments are larger ; and there seems to have been a true muscular joint be- 

 tween the second and the axillary radials, a character presented by no recent Crinoid, and 

 also, so far as I know, by no other fossil species. In Eugeniacrinus may alls from the 

 same horizon the two joints are united by syzygy, just as I believe to be the case in 

 Holopus, while the calyx is less coarsely granular. These two species are quite small 

 relatively to Holopus, the radials of Eudesicrinus not reaching a height of more than 

 1 1 mm. Associated with them in the Xe29tonct-bed (Middle to Upper Lias) of Calvados, in 

 Normandy, are some wedgershaped arm-j oints with the muscle-plates on their broad outer 

 sides produced into strong upward processes.* These joints have very much the appear- 



i Paleont. Frang., loc. cit., pi. 29, fig. 7. - Hid., pL 44, fig. 2, h. ^ Ibid, p. 78. » Ihid, pi. 11, figs. 1-4. 



