REPORT ON THE RADIOLARIA. Ixxv 



A. The group Adelacantha consists only of the suborder Actinelida, with the three 

 families Astrolophida, Litholophida, and Chiastolida (p. 728, PI. 129, figs. 1-3) ; the number of the 

 radial spines is very different and variable, sometimes only from ten to sixteen, but usually from 

 thirty to fifty, and often more than one hundred ; they are generally irregularly distributed, and 

 not as in the second main division. This latter, the Icosacantha, always possesses twenty radial 

 spines, which are regularly disposed according to a constant law, the so-called "Miillerian" or "Icosa- 

 canthan" law; the twenty spines are always so placed between the poles of a spineless axis that 

 they form five zones each of four spines ; the four spines of each zone are equidistant from each 

 other, and also from the same pole, and alternate with those of the neighbouring zones, so that the 

 whole twenty lie in four meridian planes, which cut out an angle of 45° (compare pp. 717-722, Pis. 

 130-140). In spite of the manifold variations in form which are developed in the Icosacantha, 

 they may all be derived from a common stem-form, Acanthometron (p. 742), since the law of distribu- 

 tion of the twenty spines is constantly inherited. 



B. An exception is found in the peculiar family Sphserocapsida (p. 797, Pi. 133, figs. 7—11; PI. 

 135, figs. 6-10). Here the shell is composed of innumerable small, perforated plates, which arise 

 on the surface of the calymma independently of the spines. 



III. The Skeleton of the Nassellaria. — The skeletons of the Nassellaria or Mono- 

 PYLEA consist of silica, and are never composed of separate portions, but constitute always 

 a single continuous piece. The ground-form is originally monaxon, corresponding to that 

 of the centra] capsule, with a constant difference between the two poles of the vertical 

 main axis. The ground-form is never spherical or polyaxon as in the lattice-shells of 

 the Spumellaria, and the skeleton never consists of hollow tubes as in the Ph^odaria. 

 The legion Nassellaria may be divided into two orders ; in the Plectellaria (three 

 suborders Nassoidea, Plectoidea, Stephoidea) the skeleton does not form a 

 complete lattice-shell ; in the Cyrtellaria, on the other hand, which are derived from 

 these, the siliceous skeleton forms a complete lattice-shell enclosing the central capsule. 

 The number of forms thus developed is astonishingly great, so that among the Nassellaria 

 no less than two hundred and seventy-four genera and sixteen hundred and eighty-seven 

 species may be distinguished, almost as many as in the S p h se r e 1 1 a r i a. In spite of 

 this great variety of forms the legion Monopylea is probably monophyletic ; at least all the 

 different skeletal forms may be derived from thi-ee elements which are combined in the 

 most manifold fashion ; (l) the sagittal ring, a simple siliceous ring, which lies vertically 

 in the sagittal plane of the body, encircles the central capsule and comes into contact 

 with it at the basal pole of the main axis (§ 124) ; (2) the basal or oral tripod, composed 

 of three diverging radial spines, which meet in the middle of the basal pole of the central 

 capsule (or in the centre of the porochora) (§ 125) ; (3) the cephalis, or lattice-head, a 

 simple ovoid or subspherical lattice-shell, which encloses the central capsule and stands in 

 connection with it at the basal pole of its main axis. Any one of these three important 

 structural elements of the Nassellarian skeleton may possibly be the starting-point 



