REPORT ON THE MARSUPIALIA. 103 



form and hangs freely down from the right segment of the liver behind the portal 

 fissure. The caudate lobe is comparatively speaking very small. It is prismatic in form. 



The vena cava passes through a complete tunnel of liver substance. 



The liver of the Thylacine differs very materially from that of the Cuscus. The left 

 segment is small in comparison with the right segment, and shows no tendency whatso- 

 ever to subdivision into lateral and central lobes. The right segment, however, is 

 divided into two equal portions by a well-marked cleft and the right central lobe shows 

 a deep and broad deficiency in its sharp free margin for the reception of the gall-bladder. 

 The caudate lobe is of great length (fully four inches long), and prismatic in form, whilst 

 the Spigelian lobe is small and attached by a broad base partly to the left segment, and 

 partly to the right segment of the liver. 



The vena cava is not covered in by liver substance. 



Gall-bladder and bile-ducts. — The gall-bladder in the Cuscus is very large and pyri- 

 form in shape. It is firmly attached in its upper half to the under surface of the right 

 central lobe, whilst its lower half is free and projects downwards in the cleft in the free 

 margin of this lobe. In the Thylacine the gall-bladder is very small and when distended 

 it has a globular shape ; in shape and size it is not unlike a large walnut. In both 

 animals the cystic duct emerges from the upper end of the gall-bladder, and runs upwards 

 to the portal fissure. Here it is joined by three or four hepatic ducts which issue from 

 the various lobes, and enter it by separate orifices. The common bile duct is thus formed 

 and before it opens into the duodenum it is joined by the pancreatic duct. 



The mucous membrane lining the gall-bladder of the Cuscus at first sight appears to 

 be smooth, but on close inspection it is seen to be very finely reticulated. In the 

 Thylacine it presents a very strongly marked honey-combed appearance. 



Genito-urinary Organs. 



I regret that, owing to an unfortunate accident which happened to the pelvic viscera 

 of the female Thylacine before they were examined, I am only able to give a detailed 

 account of the genito-urinary apparatus in the male Thylacine and male Cuscus. The 

 recent and very excellent paper by Dr. Young of Manchester, upon the Male Organs of 

 Generation in the Koala, 1 together with the full details given by Professor Owen in his 

 article on the Marsupialia in the Cyclopaedia of Anatomy and Physiology renders this 

 task a comparatively easy one. 



Male Thylacine (PI. X. figs. 6 and 7). 



Kidneys, ureters, and bladder. — The kidneys present the usual reniform outbne. 

 The sinus, however, is very small, and has a very constricted outlet. On section, the 



1 Journal of Anatomy and Physiology, vol. xiii. p. 305. 



