REPORT ON THE SIPHONOPHORJE. 137 



by Chun, who called it Muggisea kochii (86, p. 1157, Taf. xvii. fig. 2). Chun demon- 

 strated that this poly gastric Calyconecta is a true Monophyid, and that the cormidia, 

 arising from the common stem, become detached and sexually developed as the free 

 monogastric generation, which was described by Busch under the name Eudoxia 

 eschscholtzii. 1 Chun also gave the full description of its ontogeny, and found that the 

 larva, arising from the fertilised egg of Eudoxia eschscholtzii, does not possess the 

 pentagonal pyramidal nectophore of Muggisea, but the edgeless campanulate nectophore 

 of Monophyes ; the latter afterwards buds from the base of the former, and remains 

 when the former is detached. Chun supposed, therefore,, that three different genera- 

 tions should be distinguished in this species — (l) Monophyes pyramidalis (85, fig. 1), 

 (2) Muggisea kochii (fig. 2), and (3) Eudoxia eschscholtzii (fig. 3). I cannot agree 

 with this opinion, but I regard the first form (fig. 1) only as the larva of the 

 second (fig.. 2). The primary edgeless nectophore of Monophyes is only a provisional 

 larval organ, and the fact that it is afterwards replaced by the secondary five- 

 edged pyramidal nectophore of Muggisea may be explained by the fundamental law 

 of biogeny — by the hypothesis that Monophyes is the original ancestral form of 

 Muggisea. 



The mature Eudoxia of Muggisea is very similar to the monogastric genus Cucullus, 

 the Eudoxia of Diphyes. It differs in the rounded and edgeless surface of the conical 

 or spathiform bract, which has three or five edges in Cucullus. In respect of this 

 difference, the name Cucubalus (given in 1824 by Quoy and Gaimard, 24) may be 

 retained for it. The spathiform bract is obliquely conical, with a deep ventral groove, 

 rounded dorsal convexity, pointed apex, and simple phyllocyst (compare above, Genus 

 11b, p. 109). The free Eudoxia of the Mediterranean Muggisea kochii may, therefore, 

 bear the name Cucubalus eschscholtzii. 



A second species, slightly differing from the Mediterranean one, was observed by me 

 in the Canary Island Lanzerote, and may retain the name Muggisea pyramidalis; it 

 differs from the former mainly in the size of the conical hydrcecium, the top of which 

 attains half the length of the nectosac. The free Eudoxia of this Atlantic species has 

 a conical bract, with a blunt apex and an ovate larger phyllocyst ; it rnay be called 

 Cucubalus pyramidalis. 



A third species of Muggisea is probably the Tropical Pacific form, described by Huxley 

 in 1859 as Diphyes chamissonis. 2 It agrees with Muggisea pyramidalis in the size of 

 the high hydrceciurn, but differs from this Atlantic and from the Mediterranean species 

 in the more campanulate form of the nectophore, the denticulate shape of its edges, and 

 the stronger teeth of its mouth. The free Eudoxia of this Pacific species may, perhaps, 

 be Cucubalus cordiformis of Quoy and Gaimard. 3 Muggisea differs from the preceding 



1 67, p. 33, Taf. iv. figs. 7-10 ; Taf. v. figs. 1-9. 



2 9, p. 36, pi. i. fig. 3. 3 2 , p. 94, pi. iv. figs. 24-27 ; 24, pi. vi. fig. 1. 

 (ZOOL. CHALL. EXP. — PART LXXVII. — 1888.) Hhhh 18 



