108 



Net, som udfylder Størstedelen af Kapselen. Tab. XX. 

 Fig. 3 //. 5 g. Saavel Kapselens indvendige Side som 

 hele Netværket er beklædt med et Epithel, bestaaende af 

 smaa runde Celler, der har en rund. næsten central Kjerne, 

 Tab. XX. Fig. 3 Ti, 5 h. I Maskerne af dette Net sees 

 kortere eller længere, stavformede Legemer. Nematocyster. 

 soin i sit Indre har en fin Traad, der ikke er spiralvunden, 

 Tab. XX, Fig. 5 i, 6. Disse Nematocyster samle sig 

 efterhaanden imod Kapselens ydre Ende, hvor Nettet er 

 sparsommere og Maskerne langt større, og hvor de ligge 

 tildels pyramideformigt, med den spidse Del af Pyramiden 

 vendt mod Aabningen. Tal). XX. Fig. 5 i. Kun paa et 

 Par Papiller saa jeg Nematocyster liiige dels i Aabningen, 

 dels udenfor denne. 



Det s\nes. sum om Nematocysterne dannes og ndvik- 

 les inden disse Kapsler af Epithelialbeklædningens Celler. 



1 Maskerne sees nemlig Nematocyster i forskjellige Stor- 

 relser, og fra den runde Epithelcelle kan iagttages Over- 

 gange til forlængede Celler, der efterhaanden antage Stav- 

 form. hvori den fine Traad bliver synbar. Tab. XX, Fig. 

 5 /,-. Disse mærkelige Organer, som tor være ganske eien- 

 dommelige for Slægten Edwardsia, ihvorvel noget tilnaT- 

 melsesvis nok kan findes hos enkelte Actinider, ere visselig 

 blevne overseede af Størstedelen af de Forfattere, der have 

 beskjæftiget sig med denne Dyreslægt; kun Dr. Andres 1 

 har skjænket dem sin fulde Opmærksomhed. Han beskriver 

 dem som Nematocystbatterier uden dog at angive deres 

 indre Bygning. Naar ikke andre Forfattere have antydet 

 disse Organer, saa mener Andres, at det kan have sin 

 Grund i. at ikke alle Arter af Slægten Edwardsia ere for- 

 synede med dem; men jeg betvivler, at dette er Grunden; 

 thi hos alle de Arter, jeg har havt Anledning til at 

 observere, findes disse Papiller, dog mere eller mindre 

 fremtrædende. 



Fra den indre Kropsvæg udgaa 8 Septa, der ere 

 Forlængelser af Kropshudens Bindcvæv (Stiitzmembran, 

 La melle de soutien), og lobe fra den bagerste Ende op 

 til Mundskivens Underfiade og derfra over paa Svælgrøret, 

 paa hvilke sidste to Ste. ler de fæste sig, Tab. XX, Fig. 



2 (/. De ere meget smale i den bagre Ende. men blive 

 alt bredere og bredere, indtil de fæste sig paa Øsophagus, 

 Tal). XX, Fig. 7 a, hvorved de dele den forreste Del af 

 Gastrovascularhulheden i 8 Kamre, Tab. XX, Fig. 7 //. 

 Disse Septa, der alle ere fuldstændige, forsaavidt de fæste 

 sig paa Svælgrøret, ere ikke gjennemborede, som saa hyp- 

 pigt ere Tilfældet hos Actiniderne. saa at Kamrene ei 

 kommunieere indbyrdes med hinanden; de ere enkle, ikke 



1 Interno all' Edwardsia Claparedii (Halcampa Claparedii, 

 Panc). Memoria dell. dutt. Angelo Andresi. Mitheihmgen aus der 

 zoolog. Statiou zn tfeapel. 2 Band. pag. 123. Leipzig 1881. 



prolongations are sent forth. which anastomoze with pro- 

 longations from adjacent connective-tissue prolongations. 

 producing a reticulation , which fills the capsule for the 

 greater part (Pl. XX, fig. 3 g, 5 g). Both the internal 

 side of the capsule as well as the entire reticulation. are 

 clad with an epithelium consisting of small round cells, 

 containing a round, almost central nucleus (Pl. XX, figs. 

 3 h, 5 li). In the nieshes of this reticulation are seen, 

 shorter or longer rod-like corpuscles, nematocysts that 

 internally have a fine filament. hut which is not spirallv 

 coiled (Pl. XX, figs. 5 ?', 6). These nematocysts gradually 

 collect towards the outer extremity of the capsule, where 

 the reticulation is coarser and the meshes far larger, and 

 there they He, partly pyramidiform, with the pointed part 

 of the pyramid turned towards the aperture ■ Pl. XX. 

 fig. 5 i). Only in the case of a couple of papillæ did I 

 observe nematocysts situated partly in the aperture, partly 

 outside it. 



It appears as if the nematocysts are formed and de- 

 veloped inside those capsules, from the cells of the epithelial 

 covering. In the meshes we tind. tims. nematocysts of 

 various sizes, and from the round epithelial cells. transi- 

 tions to prolongated cells may be observed, which gradu- 

 ally assume the rod-shape in which the fine filament be- 

 comes visible (Pl. XX. fig. 5 k). Those remarkable organs, 

 which are probably quite peculiar to the genus Edwardsia, 

 althoudi a considerable resemblance to them may perhaps 

 be found in a few Actinidæ, have certainly been overlooked 

 by the greater number of writers who have occupied them- 

 selves with the studj of this animal genus; Dr. Andres ' 

 alone has devoted liis full attention to them. He describes 

 them as nematocyst-batteries without, however, indicating 

 their internal structure. The fact that other writers have 

 omitted to indicate those organs may be owing. Dr. Andres 

 thinks. to this. that not all the species of the genus 

 Edwardsia are supplied with them. hut I question whether 

 that is the reason, as in all the species that I have had 

 an opportunity of observing, those papillæ have been found, 

 although more ør less prominent. 



From the inner wall of the body 8 septa issne ; 

 these are prolongations of the connective-tissue of the in- 

 tegument of the body (stiitzmembran; Laiuelle de soutien) 

 and pass from the posterior extremity up to the interiør 

 surface of the oral disc and thence øver tø the gullet-tube, 

 upon which two last-named places they secure themselves 

 (Pl. XX, fig. '2 a). They are very narrow at the posterior 

 extremity. but become brøader and brøader until they 

 secure themselves to the cesophagus (Pl. XX. fig. 7 a), 

 dividing, thus, the anterior portion of the gastro-vascular 

 cavity info 8 chambers .Pl. XX. ti^. 7 b). These septa. 

 which are all perfect ones, in so far that they attach them- 

 selves tø the gullet-tube, are nøt perforated, as so often 



1 Interno all' Edwardsia Claparedii (Halcampa Claparedii, 

 Panc). Memoria dell. .lutt. Angelo Andresi. Mitheilungen aus der 

 zoolog. Station zu Neapel. i Band, pag. 123. Leipzig 1881. 



