16 



imedens det nedad staar i Forbindelse med det ovenfor 

 beskrevne Centralkar. I den Del af Kamret, der forlænger 

 sig ned i Stilken, ere Skillevæggene forsynede med paa- 

 tversløbende Muskelfibriller, som dog findes kun paa Sep- 

 tumets ene Side, Tab. IV, Fig. 2 c. Imedens disse Muskel-, 

 fibriller ere temmelig tydeligt at iagttage paa Skillevæggene 

 i Stilken, bar det ikke været muligt med Sikkerhed at paa- 

 vise dem længere oppe i den store, brede Del af Kammer- 

 organet, hvor Skillevæggene synes at være stærkest udvik- 

 lede. 1 Kammerorganet findes det samme Fluidum med 

 sit globulære Indhold, som ovenfor er omtalt i Centralkar- 

 ret, og saavidt jeg har kunnet erfare, sættes dette Fluidum 

 kun ved Endosmose i Forbindelse med den almindelige 

 Cirkulation. 



Det saakaldte Dorsalorgan ( kjertelformige Organ), der 

 har været Gjenstand for saa forskjellige Tydninger, og som 

 der forst i den seneste Tid er bragt nogen Klarbed orer, 

 væsentlig af Professor Perrier, har hos Bathycrinus Carpen- 

 teri ikke været saa særdeles vanskeligt at faa Rede paa, da 

 Materialet har været baade godt konserveret og nogenlunde 

 rigeligt. Dorsalorganet er beliggende i den før beskrevne 

 Centralkanal, Tab. IV. Fig. 2 d. dannet af Kammerorga- 

 nets Septa. Det er langstrakt, omgivet af en Bindevævs- 

 membran, der ligger temmelig tæt ved Kanalens Væg og 

 udvendig beklædt med Epithel af lave. næsten kubiske 

 Celler, Tab. IV, Fig. 3 a. Det er dannet af en Mængde 

 slangeformige Blindsække, adskilte ved yderst tynde Mem- 

 braner, der ere Forlængelser af den ydre Bindevævsmem- 

 brans indre Væg, hvorved det paa Tversnit faar et hippet 

 Udseende, Tab. IV, Fig. 3 b. Disse Blindsække ere iml- 

 vendig beklædte med et Epithel. der bestaar af temmelig 

 store, runde Celler med Kjerne og Kjernelegeme, og hvis 

 Indhold er et yderst fint kornet Protoplasma. Tab. IV. Fig. 

 3 c. Naar Dorsalorganet er kommet ovenfor det femkara- 

 rede Organ, lægger det sig lidt til Siden imellem to Tarm- 

 slynger, Tab. IV. Fig. 3 d. 4 a. og stiger nu op mod Mund- 

 skiven i Hoide med Forbindelsen imellem 2det og 3die 

 Radiale, hvor det afgiver 5 Stammer, som synes at være 

 forbundne med Tvergrene, Tab. IV, Fig. 4 b. Hver af 

 disse 5 Stammer dele sig ved den overste Del af det 3die 

 Radiale i to Grene, der gaa til hver sin Arm, Tab. IV, 

 Fig. 4 c; hver Gren fortsætter nu Lobet i Genitalkanalen, 

 som ligger i Skillevæggen imellem Dorsal- og Ventralkana- 

 len, og forlænger sig ind i Pinnulen, hvor den udvider sig 

 noget og danner her det egentlige Kjønsorgan. 



Saavel de 5 Stammer som senere deres to Grene er 

 paa deres indre Væg, altsaa ogsaa paa det indre Genital- 

 rors Væg, beklædt med et lignende Epithel, som det. der 

 er beskrevet ovenfor ved det kjertelformige Organ (Dorsal- 

 organet). Det indre Genitalrørs ydre Væg ligger ganske 

 til det ydre Genitalrørs indre Væg, saa at det er yderst 



any aperture that could place it in direct eonnection with 

 the irrigation apparatus described by Professor Perrier. 

 whilst at the foot it is placed in eonnection with the 

 central vessel described above. ln that portion of the 

 chamber that prolongs itself down in the stalk. the divis- 

 ional walls are furnished with obliquely running museular 

 fibres, which are, however, only found on the one siile of 

 the septum (Pl. IV, fig. '2 c). Whilst these museular fibres 

 may be pretty distinctly observed on the divisional walls 

 in the stalk. it has not been possible to discover them, 

 with certainty, farther up in the large, broad portion of 

 the chambered organ, where the divisional walls appear to 

 be most developed. The same fluid with its globular con- 

 tents, as is spoken of previously as contained in the cen- 

 tral vessel, is found in the chambered organ, and, so far 

 as I have been able to expeuence, this fluid is placed in 

 eonnection with the general cireulation only by endosmose. 



The so-called dorsal organ (gland-shaped organ) which 

 has been the object of sucb different explanations. and in 

 regard to which tliere first in quite late times lias been 

 thrown some light, chietly by Professor Perrier. has ;is 

 regards Bathycrinus carpenteri not been so very difficult 

 to elucidate, as the material has been both somewhat 

 abundant and well preserved. The dorsal organ is situated 

 in the first described central canal (Pl. IV, fig. 2 cl) formed 

 by the septa of the chambered organ. It is elongate. and 

 surrounded by a connective-t issne membrane that lies 

 pretty dose to the wall of the canal, and is exteriorly clad 

 with an epithelium of low, almost cubical cells (Pl. IV, fig. 

 3 a). It is formed by a multitude of anguilliform ca^ca 

 separatéd by extremelv thin memjjranes, which are prolon- 

 gations of the inner wall of the exterior connective-tissue 

 membrane. whereby. a patched appearance is imparted to 

 transversal sections (Pl. IV, fig. 3 b). Those cæca are clad 

 internallv with an epithelium consisting of pretty large. 

 round cells with nucleus and corpuscle, and containing au 

 extremelv finely granulated protoplasm (Pl. IV. fig. 3c). 

 When the dorsal organ has arrived above the five-chamb- 

 ered organ, it lavs itself a little to the side between two 

 iutestinal twistings (Pl. IV, fig. 3 d. 4 a). and then rises 

 up towards the oral disc to a level with the eonnection 

 between the 2nd and 3rd radials, where it gives off 5 

 stems which appear to be connected by cross-branches 

 (Pl. IV. fig. 4 b). Each of those 5 stems divides itself at 

 the uppermost part of the 3rd radial into two branches, 

 each of which passes over to an arm (Pl. IV. fig. 4 c); 

 each branch then continues its course in the genital canal, 

 which is situated in the divisional wall between the dorsal 

 and ventral canal, and prolongs itself into the pinnule, 

 where it expands itself somewhat and forms here the real 

 sexual organ. 



The 5 stems as well as, subsequently. their two 

 branches are, upon their inner wall. consequently also on 

 the inner wall of the genital tube, clad with an epithelium 

 similar to that described above in eonnection with the 

 gland-shaped organ (the dorsal organ). The outer w r all of 

 the inner genital tube lies quite in to the inner wall of 



