ACTINIARIA 



21 



Fam. Halcampoididae 



Fam. Edwardsiidae Subfatn. Edwardsiinae Genera. Edwardsia Quatr. 



Isoedwardsia Carlgr. 

 Subfam. Milne-edwardsiinae . . . Genera. Milne-edwardsia Carlgr. 



Paraedwardsia Carlgr. 

 Genera. Halcampoides Dan., Acthelmis Liitk., Phytocoetes- Ann., Halcampclla 

 Andr., Synhalcampella^ Carlgr., Scytophorus R. Hertw., Pentactinia 

 Carlgr., Harenaciis Torr., Siphonactinopsis Carlgr. Mesacmaea Andr., 

 Peachia Gosse, Eloaclis Andr., Haloclava Verr., PPolyopis R. Hertw. 

 Genera. Limnactinia Carlgr., PPolyopis R. Hertw. 



Genera. Halcampa Gosse, Parahalcampa Carlgr., Synhalcampa- Carlgr., Cacto- 

 soma Dan., Mena'^ Steph., Halianthella Kwietn. 

 Fam. Halcampactiidae. Genera. Halcampactis 'Parquh., Haliactis Cailgi. , Pelocoetes Ann.^, Pllyactis Andi., 



POctophdlia Andr. 

 Fam. Andwakiidae. Genera. Andwakia Dan., Octineon Mosel, Pllyactis Andr., POdophellia Andr. 



Fam. Ilyanthidae. Genera. Ilyanthus Forb., Oractis Mc. Murr. 



Fam. Limuactiniidae 

 Fam. Halcampidae. 



In the following I will further discuss the position of the genera within the families of which I have 

 mentioned the first six here. Concerning the position of the genus Oractis I am a Uttle doubtful. It is true 

 that, according to me, it cannot be placed in the family Gonactiniidae, as proposed byMc. Murrich, because 

 it is only in the arrangement of the mesenteries that it partly agrees with this familj', while it differs essenti- 

 ally from it in the other characters. The only families which are to be considered in the placing of this genus 

 are the Halcampoididae and the Ilyanthidae. The endodermal sphincter which is, in proportion to the small 



1 Compare the family Halcampoididae. 



^ Since this was written Stephenson (1920 p. 520) has proposed a new family provided with acontia, Diadumcuidae, 

 enclosing Diadumene Ann, Pelocoetes Ann., Phytocoetes Ann., and Mena Steph. The family is ccrtainl)' heterogeneous. The type- 

 genus differs in several respects from the three others. The former has a well-developed pedal disc and certainly belongs to 

 the BasUaria, while the three latter probablj' are Abasilaria. (It is probably a lapsus of Annandale when he (1915 p. 81) speaks of 

 basilar muscles instead of parietal rau.scles). In consequence of .A.nnandale's description of their structure it seems to me that 

 Pelocoetes would belong to the Halcampactiidae, Phytocoetes and Mena to the .\ndwakiidae. But on fiu-ther examination of .An- 

 nandale's figure 3 (p. 80) in which the acontia are also represented, I think that at least Phytocoetes gangicus is not provided 

 with acontia. The figure designated as acontia is namely no such thing, but simply mesenterial filaments. Thus Phytocoetes is a 

 Halcampoid and nearly related to Acthelmis. Concerning Mena {Phytocoetes) c/n7Aaea I am of opinion that this species is a Cac/o- 

 soma. Nothing in the structure speaks against it ; on the contrary the appearance of the sphincter and the presence of papillae on 

 the column indicate that we have to do with this genus. At least it is closely related to this genus. Finally Pelocoetes is a good genus 

 and probably belonging to the Halcampactiidae. Though .\nuandale has not given any figure of the acontia, but says that they are 

 long and relatively stout, it is probable that in this case he has not mistaken the mesenterial filaments for acontia. 



The species Halianthus limnicola Ann. is, though a Halcampid, probably the type of a new genus. It is neither a Hal- 

 campa nor a Cactosoma. The absence of secondary, imperfect mesenteries in the whole length or almost so of the column and the pres- 

 ence of 12 rows of tubercles on the coliunn, ser\'e to distinguish it from the two genera. Besides, it differs from Halcampa in having 

 sometimes more than 12 tentacles, but in this respect it agrees with Caftoso»H«. The presence of extra-tentacles indicates that there 

 are weak mesenteries in the uppermost part of the coluum as in Halcampclla of the Halcampoididae. As Halianthus is a synonym 

 of Halcampa I propose a new name for it Synhalcampa, characterized as follows: 



Halcampidae with no external differentiation of capituluni, scapus and physa. .\boral extremity provided with a porus. 

 Column with 12 longitudinal rows of solid tubercles, towards the aboral end obsolete. Sphincter weak, close below the base of the 

 tentacles. Tentacles 12 or some more, short, but stout and cyhndrical. Two rather well-developed siphonoglyphes and 2 pairs of direct- 

 ives. Six pairs of perfect mesenteries with rather strong pennons. When the tentacles are more than 12, some very weak mesenteries 

 are probably found in the uppermost part of the column. Excepting these, no imperfect mesenteries. 



