ACTINIARIA 



59 



Subfam. Milne-Edwardsiinae. 



Diagnosis: Edwardsiidae without neniathybomes in the scapus. Nematocysts in the ectoderm 

 of the scapus scattered or in heaps. Physa absent, indistinct or feebly developed. Nematocysts of the capi- 

 tulum of almost the same size as those of the scapus. Inner tentacles longer than the outer ones ; commonly 

 hexamerously arranged. 



This subfamily corresponds to the family Milne-Edwardsiidae, proposed by myself (1893, p. 11), 

 and is easily distinguished from the subfamily Edwardsiinae by the absence of nemathybomes. Also the 

 arrangement of the tentacles is another one, at least in the genus Milne-Edwardsia, but probably also in 

 Paraedwardsia as I have been able to state the same arrangement in P. sarsii as in Milne-Edwardsia. The 

 tentacles are namely commonly hexamerously arranged, and the inner tentacles are larger than the outer 

 ones (textfig. 11), so this is another agroupment than that of the genus Edwardsia. Concerning the num- 

 ber of siphonoglyphes I can ascertain that several species have only one, a ventral one. Whether this is 

 characteristic of the whole subfamily, remains to be confirmed. 



This subfamily does not seem as rich in species as the subfamily Edwarsiinae. Very likely the num- 

 ber of Milne-Edwardsia-s-pecies will increase, when all Edwardsiidae have been subject to a more detailed 

 examination. There is no doubt that several species, described as Edwardsiella and Edwardsia, belong to 

 this subfamily. Thus, according to my examination, the Edwardsia timida Quatr. is a Milne-Edwardsia, 

 M.dixonii, (verified by myself on material received from Dixon). Besides this, it is not improbable that one 

 part of the forms, described and reproduced by Andres 1883 as varieties of Edwardsia claparedii, are in 

 fact Milne-Edwardsia- or Paraedwardsia-s^eci&s. In the subfamily I included two genera Milne-Edwardsia 

 and Paraedwardsia, of which the latter is furnished with "i7a/cflw/)«-papillae", the former not. 



Genus Milne-Edwardsia Caiigr. 



Diagnosis: Milne-Edwardsiinae with the column divisible into a lower, greater part which is 

 invested with a rather well-developed, sometimes very thick cuticle and an upper minute part, capitulum, 

 without cuticle. A weak physa also sometimes present. Scapus without nemathybomes and " Halcampa-papUlsie" . 

 Nematocysts in the ectoderm of the scapus either scattered or arranged in groups, comparatively short, in 

 proportion to the breadth. Nematocysts of the capitular ectoderm commonly large and mainly distributed 

 on the ridges of the capitulum. Capitulum more or less polygonal. Tentacles 12 or in several cycles, hexa- 

 merously arranged, the inner longer than the outer ones (alvays?). Only one, ventral siphonoglyphe (always?). 



In a certain respect the diagnosis of this genus, pubhshed in this paper, is a httle more exphcit 

 than the original one. The hitherto known species of this genus are: M. loveni Carlgr., M. carnea (Gosse), 

 M. polaris Carlgr., M. nathorstii Carlgr. and M. dixonii Carlgr. (nov. nomen pro "Edwardsia timida Quatr." 

 described by Dixon). It is also possible that Edwardsia lineata Verr. will be included in this genus. The 

 species described below are easily distinguished from each other. 



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