233 



THE GENITALIA OF THE TSETSE FLY, GLOSSIMA 



PA LP A US. 



By W. Wesche, F.RM.S. 



(Bead Jvne loth, 1905.) 



Plate 15. 



While carrying out some dissections of the mouth-parts of 

 Glossina palpalis Des., the host of the germ of sleeping sickness, 

 the genitalia attracted my attention owing to their large size and 

 great complexity. I find that they homologise fairly well with 

 those of Scatophaga, which I described in 1903*; but the com- 

 plexity of the central organ — the penis, adminiculum, or appendix 

 interna — has necessitated the addition of fresh terminology to 

 that used in the above-mentioned paper. 



The males in Diptera may be divided into two classes: (1) 

 those which have the segment containing the genitalia (the 

 hypopygium) turned in under the abdomen; and (2) those in 

 which this segment forms the end of the abdomen, the penis and 

 appendages being sunk in a cavity, and the outer claspers or 

 forcipes superiores quite evident at the extremity. These latter 

 are of endless variety both in shape and size ; they are very 

 prominent in the families Tipulidae, Culicidae, Cordiluridae, and 

 Anthomyidae, and I propose to classify these as " hamate." 



In the Muscidae, including the families Sarcophagidae and 

 Tachinidae, the Syrphidae, as well as the smaller families of 

 Conopidae, Pipunculidae, Lonchopteridae, and Phoridae, the 

 hypopygium, armed as in the hamate families with the forcipes 

 superiores on the ultimate segment, is turned in under the abdomen, 

 hiding the claspers; this division I propose to call " sub -hamate." 

 In this must be included the Dolichopidae, or Fan-tailed Flies, 



* Journal of the Quekett Microscopical Club, Series 2, Vol. 8, No. 52. 



