Messrs. Hancock and Embleton on the Anatomy of Eolis. 85 



animal is moving freely about ; and until this be done, all theo- 

 rizing appears to us idle, and likely to lead to error and confu- [ 

 sion. 



Salivary glands. — These in Eolis are very small and difficult 

 to detect; they lie concealed between the corneous plates and 

 the muscular mass of the cheek, as previously noticed. On 

 removing either of the corneous plates in E. papulosa^ the gland 

 will be found lying exposed in a depression on the upper third 

 of the external surface of the cheek-mass, corresponding to about 

 the centre of the corneous plate, PI. III. fig. 6 a. The gland is 

 composed of a small cluster of roundish, yellow, irregular folli- 

 cles, fig. 5, and frequently a little way in advance of this there 

 is a smaller one made up of two or three follicles. The two parts 

 are connected by a long slender duct, which passing backwards 

 opens into the mouth at the commencement of the oesophagus. 

 We have likewise detected this gland in E. coronata. 



This gland differs conspicuously in size, position and character 

 from the same organ in Eolidina, figured and described by M. de 

 Quatrefages. All we can say is, that in our researches we have 

 observed nothing like the representation given by him of the 

 salivary glands of that species. It is certainly not likely that in 

 animals so closely allied these organs should be so widely different. 

 We would suggest therefore the possibility of that naturalist 

 having mistaken some portion of the generative organs for them. 

 We are inclined to do this the more since he has entirely mis- 

 understood the sexual apparatus, and figured and described only 

 a small portion of it, and since we have sometimes observed, when 

 examining these animals in the compressor, portions of that ap- 

 paratus not altogether unlike M. de Quatrefages^ figure of the 

 salivary glands of Eolidina paradoxum. 



Since wiiting the above, we have had much satisfaction in 

 gathering from the Observations of M. Souleyet on the Gaste- , 

 ropod Mollusca, forming the proposed order Phlehenterata of M. 

 de Quatrefages, translated in the November Number of the 'An- 

 nals,' that our views have been almost altogether verified. 



With respect however to the anatomy of the gland of the pa- 

 pillse, M. Souleyet appears to adopt the opinion of M. de Qua- 

 trefages, w^ho states that the prolongation of the digestive cavity 

 into the papilla is coated with a granular layer — the liver. We 

 have shown above that this view is inaccurate, and in confirma- 

 tion of this we may as well state, that on the papillae being sub- 

 jected to pressure, the granular structure of the gland invariably 

 becomes ruptured internally; but if the view taken by these 

 gentlemen be correct, we should apprehend that the rupture 

 would take place externally into the vascular canal surrounding 

 the gland. 



