A revision of the genus Anacyclus L. (Compositae :\^ 

 Anthemideae) 



Christopher John Humphries 



~ k 



Department of Botany, British Museum (Natural History), Cromwell Road, London SW7 5BD 



Contents 



Synopsis ............. 84 



Introduction ............. 84 



Materials and methods ........... 84 



Descriptive terminology .......... 84 



Taxonomic concepts ........... 85 



Historical outline ............ 86 



Delimitation and systematic position ......... 89 



Morphology ............. 91 



Habit 91 



Leaves ............. 91 



Inflorescence ............ 93 



Receptacle ............. 95 



Involucre ............. 95 



Florets 95 



Cypselas 96 



Chromosome numbers .......... 98 



Phytogeography 101 



Phylogeny 102 



Systematic descriptions ........... 109 



Anacyclus L. ............. 109 



Artificial key to species, subspecies, varieties and hybrids 110 



Sect. 1- Pyrethraria DC Ill 



1. A. pyrethrum (L.) Link . . . . . . . . .111 



a. var. pyrethrum . . . . . . . . . .114 



b. var. depressus (Ball) Maire . . . . . . . .114 



Anacyclus pyrethrum in medicine and the origin of A . officinarum Hayne . 114 



Sect. 2. Anacyclus . . . . . . . . . . .116 



2. A. monanthos (L.) Thell 116 



a. subsp. monanthos . . . . . . . . .118 



b. subsp. cyrtolepidioides (Pomel) Humphries . . . . .118 



3. A. maroccanus (Ball) Ball 119 



4. A. radiatus Loisel 120 



a. subsp. radiatus . . . . . . . . . .123 



b. subsp. coronatus (Murb.) Humphries . . . . . .124 



5. A. clavatus (Desf.) Pers 124 



6. A. homogamos (Maire) Humphries ....... 127 



a. A. x inconstans Pomel . . . . . . . .130 



b. A. x valentinus L. ......... 131 



7. A. linearilobus Boiss. & Reuter . . . . . . . .131 



8. A. latealatus Hub.-Mor 134 



9. A. nigellifolius Boiss. 135 



Excluded taxa . . . . . . . . . . . .137 



Acknowledgements . . . . . . . . . . . .137 



References ............. 137 



Taxonomic index . .140 



Bull. Br. Mus. not. Hist. (Boc.) 7 (3) : 83-142 Issued 20 December 1979 



83 



