REVISION OF ANACYCLUS 

 Table I (cont.) 



105 



Character 



Plesiotypic ( ) 



Character state 



Apotypic ( + ) 



Leaves 



1 7 Leaf rosette 



1 8 Leaves 



1 9 Leaves 



20 Sinus between 2nd-order 

 lobes 



21 Basal leaf divisions 



22 Basal leaf divisions 



23 Leaves 



24 Terminal leaf-lobes 



Capitulum 



25 Capitulum 



26 Ligule colour 



27 Ligules 



28 Disc corollas 



29 Disc corollas 



30 Disc corollas 



31 Disc corolla articulation 



32 Disc corolla wings 



33 Wings 



34 Receptacular scales 



35 Receptacular scales 



36 Receptacular scales 



37 Scales 



38 Involucral bracts 



39 Involucral bracts 



40 Involucral bract margins 



41 Inner involucral bracts 



42 Cypselas 



43 Cypsela wings 



44 Cypsela wings 



45 Cypsela wings 



46 Cypsela auricles 



47 Cypsela development 



48 Pappus 



absent 



homomorphic 



herbaceous 



narrow, less than 2 cm 



many 



tripinnatisect 

 vermiform 

 linear-lanceolate 



radiate 



white above/red below 



exerted above involucral 



bracts 



actinomorphic 



articulate 



articulate 



terminal 



wingless 



narrow, usually parallel 



membranous 



unthickened 



oblanceolate 



erect at apex 



herbaceous 



obtuse 



fuscate 



herbaceous 



terete 

 absent 



margins smooth 

 margins smooth 

 absent 



terete or rhomboid during 



development 



absent 



present 



heteromorphic 



fleshy 



wide, more than 2 cm 



few 



bipinnatisect 



bi- or tripinnatisect 



oblanceolate 



discoid 



yellow above/red below, 



'yellow, "white 



inserted below the involucral 



bracts 



zygomorphic 



anteriorly vaginate 



posteriorly vaginate 



anterior 



winged 



large, oval 



tough 



dorso/anteriorly thickened 



lanceolate 



overlapping at apex 



papery 



acuminate or long-pointed 



light brown/hyaline, colourless 



expanded, membranous and 



hyaline at the apex 



anterior-dorsally compressed 



narrow/thick 



'narrow/thin 



" broad/thinner 



margin crispate 



margin spiny 



thick/pointed 



' thin/pointed 



" thin/rounded 



flat throughout development 



marginal, 



' marginal coroniform 



" coroniform 



By using Hennig's (1965, 1966) methods, it has been possible to reconstruct a phylogenetic 

 cladogram (Fig. 10) which provides the most parsimonious evolutionary model to account for 

 morphological variation within this group and a basis for the interpretation of cytogenetic and 

 biochemical data (Ehrendorfer et al., 1977; Humphries in press a). The sister group of Anacyclus 

 and Leucocyclus is indicated by inclusion in the diagram of other genera within the Anthemideae 

 assemblage - a group taken to comprise Achillea and its allies. 



