REVISION OF ANACYCLUS 



VAL 



107 



/ 



PYR 



-19,20,24,39- 



LA NIG 



-26.31', 40,46 



8* .10.1 2.14, 16 ,25* t 34*.35,(45)- 



28. 31. 32. 43 1 ,48 



-(2), 4. 5. 8*. 11, 13.17,18,34*37 - D 



1.3,7 



Fig. 10 Cladogram showing phylogenetic relationships in the Anacyclus group, based on morpho- 

 logical characters in Tables 1 & 2 and reconstructed according to the principles of Hennig: AN - 

 Anthemideae (Achillea and its allies), PYR - A. pyrethrum, MO - A. monanthos, OFF - A. 

 officinarum, MA - A. marrocanus, RAD - A.radiatus, VAL - A. X valentinus, CLA - A. clavatus, 

 INC - A. x inconstans, HOM - A. homogamos, LIN - A. linearilobus, LA - A. latealatus, NIG - 

 A. nigellifolius, LEU - Leucocyclus formosus. 



Hennig's method makes it necessary to interpret character states either as primitive or derived 

 conditions. This has been carried out by the principle of sister-group or related-group comparison 

 so that commentary on some of these plesiotypic or apotypic features will help to explain the 

 rationale behind the reconstruction of species relationships within Anacyclus. 



In general, the systematic distribution of the majority of character states considered to be 

 derived is consistent with the status accorded them here. Thus, for example, the apomorphic 

 conditions found in characters 5, 10, 13, 14, 29, 30, 42, 43, 46 and 47 (Table 1) represent structural 

 modifications which appear to be absent from all groups except those which they help to define. 

 Therefore, Anacyclus and Leucocyclus form a monophyletic assemblage by the unique possession 

 of the character states 42 and 47 (Tables 1 & 2). 



