REVISION OF ANACYCLVS 125 



Anthemis pubescens Willd., Sp. PI. 3(3) : 2177 (1800). Orig. coll.: Herb. Willd. 16246 (B, holotypus). 

 Anthemis biaristata DC. in Lam., Fl. Fr. 4 : 204 (1805); Biv. PI. Sic. Cent. 2 : 1 (1807). Orig. coll.: 



Herb. DC., 'biaristata, les champs N.' (G-DC, holotypus). 



Anacyclus pedunculatus (Desf.) Pers., Syn. PI. 2 : (1807), pro par te quoad basionym. 

 Chamaemelum inodorum sensu Cup., Pamph. 2 : t. 69 (1807), non Vis. 

 Anacyclus divaricatus Cav. ex Balbis Cat. Hort. Taur. : 11 (1813); Steudel, Nomencl. Bot. 1st ed. : 41 



(1821), nom. nud. 



Anacyclus tomentosus sensu DC. in Lam., Fl. Fr. 5 : 481 (1815). 

 Chamaemelum incrassatum Hoffmanns. & Link, Fl.Port. 2 : 348 (1820). Orig. coll.: 'Frequent sur le 



bord de la riviere du Nabao pres de Thomar' ( ? typus destructus). 

 Anthemis incrassata (Hoffmanns. & Link) Link, Enum. Hort. Berol. Alt. 2 : 345 (1822). 

 Anacyclus pubescens (Willd.) Reichb., Fl. Germ. Excurs. 2 : 225 (1831). 

 Bambagella clavata (Desf.) Ten., FL Nap. 5 : 235 (1835). 



Anacyclus mucronulatus Hort. ex Steudel, Nomencl. Bot., 2nd ed. : 82 (1840) nom. nud., non Guss. 

 Anacyclus tomentosus var. P marginatus Guss., Fl. Sic. Syn. 2(1); 495 (1844), nom. illeg. 

 Anacyclus candolii Nyman, Syll. : 8 (1854). 



Anacyclus aristulatus Link ex Nyman, Consp. 2 : 363 (1879), nom. nud. 

 Anacyclus clavatus var. a typicus Fiori & Paol., Fl. Anal. Ital. 3 : 261 (1904), nom. illeg. 

 Anacyclus clavatus var. P tomentosus (L.) Fiori & Paol., Fl. Anal. Ital. 3 : 261 (1904). 

 Anacyclus tomentosus forma glabrus [sic] Huter, Porta & Rigo in Fiori, Fl. Anal. Ital. 3 : 261 (1904), 



nom. nud. et illeg. 

 Anacyclus capillifolius Maire in Bull. Soc. Hist. nat. Afr. N. 22 : 296 (1931). Orig. coll: Font Quer & 



Maire s.n. (25 June 1930) 'In Atlante rifano: in lapidosis calcareis montis Kraa, 1600m' (MPU, 



holotypus). 



Illustrations: Figs 2, 4, 19. Reichenbach, 1854 : tab. 999, fig. II, 4-11 ; Cesati, Passer & Gibelli, 1867-86; 

 tab. 79, fig. 5a; Cusin & Ansberque, 1873 : tab. 146; Hoffman, 1894 : 269 fig. K; Acloque, 1894 : 374; 

 Fiori & Paoletti, 1904 : tab. 426, fig. 3590; Merino, 1906 : 365; Lazaro e Ibiza, 1907 : 658; Adamovic, 



1911 

 1922 

 1976 



tab. 66 : Cadevall & Salient, 1917 : p. 277, fig. 1421 ; Small, 1918 : p. 22, fig. K, M. : Bonnier, 

 tab. 297, fig. 1473 : Ponzo, 1927 : p. 563, fig. 24; Negre, 1962 : p. 285, fig. 712 a-c: Zhangeri, 

 tab. 143 : figs 5238, 5241; Heywood & Humphries, 1977 : p. 865, fig. 7. 



Common names: Rebiana, Beehibchou, Bouibicha, Bo Melal, Math-el-Djadja, Redjelet el Rh'orab, 

 Oum-el-ali, White Anacyclus. 



Annual; stems 7-0-30-0 (-40-0) cm, slender to stout, procumbent to erect, usually much branched 

 from the base or branching from the middle, sparsely to densely appressed villous. Leaves 

 (1-5-) 2-5-11 -5 x(0-5-) -3-6 cm tri- to bipinnatisect, oblanceolate in outline, glabrescent to 

 densely villous, usually sessile but occasionally long-petiolate; primary lobes in 3-12 subopposite 

 or opposite pairs; ultimate segments linear-lanceolate 3-0-7-0 x c. 0-5 mm; rhachis flat, cuneate 

 at the base with 3 or more prominent veins. Inflorescence a lax, corymbose cyme with up to 40 

 erect or ascending peduncles; peduncles markedly clavate near the apex at maturity, villous 

 just below the capitulum. Involucre (0-5-) 8-0-1-8 mm in diameter, hemispherical; involucral 

 bracts in three rows, 1 -8-7-0 x 0-5-2-0 mm, outer series linear-triangular, inner series rectangular 

 usually densely villous, pale brown or green and thicker towards the centre with thin, scarious, 

 entire, pale yellow to dark brown margins; receptacular scales 1 -7-5-0 xc. 2-5 mm, obovate or 

 cuneate and mucronate, slightly caniculate towards the apex, somewhat membranous. Ray 

 florets white; ligules dimorphic, 5-5-16-5 x 2-0-7-0 mm, white, trifid, usually somewhat rounded, 

 creamy white; tube 3-5-5-5 x 0-5-1-0 mm, anterior-dorsally compressed with narrow, + parallel- 

 sided wings. Disc florets 3-5-5-0x0-5-1 -3 mm, hypocrateriform with a distinct, campanulate 

 corolla and a narrow basal tube; tubes anterior-dorsally compressed with broad, rounded wings 

 towards the base, up to 1-0 mm wide in outer series, but narrower, c. 0-6 mm wide, towards the 

 centre of the disc; lobes usually dimorphic, with three short lobes 0-5-0-6 mm long and two 

 extended lobes, up to 2-0 mm long, especially towards the centre of the disc. Styles 3-0-4-0 mm 

 long, style branches 0-3-0-6 mm long. Stamens c. 4-0 mm long, anthers 1-8-2-0 mm long. Cypselas 

 (1-5-) 2-5-3-5 (-5-0) x (0-4) 1-5-4-7 mm, broadly obovate, those of the ray and outer disc with 



