128 C. J. HUMPHRIES 



Tahanout ad radices Atlantis majoris it ditione Reraya, ad alt 900-1000 m' (RAB, holotypus, MPU, 



isotypus). 

 Anacyclus tomentosus (var.) c. discoideus Guss., Fl. Sic. Syn. : 495 (1844). Orig. coll.: Gussone s.n., 



Italia Caltanisetta, Lentini Alicata (NAP, lectotypus) (not seen). 

 Anacyclus clavatus (var.) P discoideus Willk. & Lange, Prodr. Fl. Hisp. 2 : 84 (1865); Batt. & Trabut, 



Fl. Alger. : 452 (1890), pro pane. 



Anacyclus valentinus sensu Briquet & Cavallier in Burnat, Fl. Alp. Marit. 6 : 165 (1916), non L. 

 Anacyclus dissimilis var. australis Maire in Bull. Soc. Hist. not. Afr. N. 20 : 186 (1929) Orig. coll. : Maire 



654, 'Hab in alveis arenoso-limosis torrentium in montibus Hoggar. Saharae centralis, ad alt. 



1200-1450 m' (AL, lectotypus, MPU). 

 Anacyclus valentinus subsp. eu-valentinus Thell. in Jahandiez & Maire, Cat. PI. Maroc 3 : 767 (1934), 



nom. nud. 

 Anacyclus valentinus subsp. dissimilis sensu Negre, FL Maroc Aride 2 : 284 (1962), quoad descr. 



Illustrations: Figs 2, 4, 21. Gaertner, 1791 : tab. 165; Lamarck, 1798 : tab. 700. fig. 1; Jaume St-Hilaire, 

 1808 : tab. 27; Negre, 1962 : tab. 124, fig. 713. 



Common names'- Guerthoufa. 



Annual; stems 5-0-40-0 (-50-0) cm, slender, ascending to erect, usually branched from the base, 

 sparsely to densely appressed villous. Leaves ( 1 -0-) 4-0-9-0 (-13-0) x 0-3-4-0 cm. tri- to bipinnatisect 

 oblong or oblanceolate to obovate in outline, usually hairy, sessile. Inflorescence a lax corymb, 

 with erect or ascending peduncles, emerging from a short stem; peduncles usually clavate at 

 maturity, usually villous just below the capitulum. Involucre 5-0-1 8-0 mm in diameter, hemis- 

 pherical; involucral bracts multiseriate, 3-0-7-0 x 1-8-3-0 mm; outer series linear-triangular, 

 inner series rectangular, usually villous towards the centre and apex, brown or pale greenish with 

 a thin brown scarious margin. Receptacular scales 3-0-4-5 x 1-1-1-7 mm, obovate or cuneate and 

 mucronate, slightly caniculate towards the apex, somewhat membranous. Capitula discoid, all 

 florets monoecious discoid. Florets 2-5-4-6 mm long, hypocrateriform with a distinct campa- 

 nulate upper part; tube slender, anterior-dorsally compressed with broad, rounded wings towards 

 the base; wings up to 1-2 mm wide in the outer series, c. 0-7 mm wide towards the centre of the 

 capitulum, lobes irregularly dimorphic, particularly in central florets with three short lobes, 

 0-3-0-9 mm long and 2 long lobes up to 1-6 mm long. Styles c. 4-0 mm long, style branches 

 0-2-0-5 mm long. Stamens 3-8-4-0 mm long, anthers 1-2 mm long. Cypselas 1-5-4-2 x 1-4-3-8 mm, 

 broadly ovate, with thin membranous wings; wings somewhat heteromorphic, those of the outer 

 series with broad wings 0-7-1-8 mm wide with rounded or slightly pointed diverging auricles, 

 those of the inner series with narrow wings 0-4-0-6 mm wide and less prominent auricles. Pappus 

 a fimbriate corona contiguous with the wings, particularly developed on the anterior side of the 

 apex. 



Flowering period: March- July. 

 Chromosome number: 2n= 18. 



Observations: This species is easily recognized by its broad, more or less parallel-sided, cypsela 

 wings and the somewhat rounded wings at the base of the corolla tube. It is very closely related to 

 Anacyclus clavatus and is considered in this revision to be its sister-species (see p. 108). However, 

 it is a much less robust species, the capitula are obviously discoid with homogamous herma- 

 phrodite disc-florets, and the cypselas have more delicate, but broader wings. 



Many apparently discoid specimens of Anacyclus have been identified as A. valentinus L. when 

 in fact they are not homogamous discoid but heterogamous with structurally reduced female 

 ligules in the outer floret series of the capitulum (Fig. 19). There seem to be two possible reasons 

 for this phenomenon: (i) either the short-liguled forms represent an intermediate stage in the 

 evolution of discoid monoecious forms from gynomonoecious radiate taxa or (ii) the short- 

 liguled plants are hybrids between discoid and radiate taxa. 



