REVISION OF ANACYCLUS 131 



b. Anacyclus X valentinus L. 



Sp. PL : 892 (1753) (A. homogamosx radiatus). (Lectotypus, see p. 109). 

 Anacyclus lanuginosus Moench., Meth. : 581 (1794) Orig. coll.: typus destructus. 

 Anacyclus clavatus (var.) P discoideus Willk. & Lange, Prodr. Fl. Hisp. 2 : 84 (1 865), propane. 

 Anacyclus valentinus (var.) 3 microcephalus Costa in Willk. & Lange, Prodr. Fl. Hisp. 2 : 84 (1865); 



Sennen in Sennen & Mauricio, Cat. Fl. Rif. Or. : 59 (1933). Orig. coll.: Costa PL catal exs. (M, 



holotypus) (not seen). 

 Anacyclus dissimilis Pomel, Nouv. FL Atl. : 53 (1874). Orig. coll.: Pomel s.n. (1 April 1862),Terrains 



Sablonneux du Sahara, Mzab, Metlili (Al, holotypus, MPU). 

 Anacyclus prostratus Pomel, Nouv. Mat. FL Atl. : 52-53 (1874). Orig. coll.: Pomel s.n., 'Lieux herbeux 



des montagnes elevees d 1'interieur (Al, holotypus) (not seen). 

 Anacyclus radiatus (var.) 3 valentinus (L.) Arcangeli, Comp. FL Ital. : 359 (1882). 

 Anacyclus radiatus (var.) 3 discoideus Chiov. in Fieri & Paol., FL Anal. Ital. 3 : 261 (1903) Orig. coll.: 



Chiovenda, 'Lit. del Lazio tra Furbana e S. Severa (Fl, holotypus) (not seen). 

 Anacyclus valentinus var. ligulata Sennen in Anvari, Jta Cienc. Nat. Bare. 2 : 647 (1917), nom. nud. 

 Anacyclus valentinus subsp. dissimilis var. typicus Maire in Bull. Soc. Hist. not. Afr. N. 20 : 186 (1929), 



nom. illeg. 



Anacyclus valentinus var. eriolepis Maire in Bull. Soc. Hist. nat. Afr. N. 20 : 186 (1929), nom super fl. 

 Anacyclus valentinus subsp. dissimilis (Pomel) Thell. in Jahandiez & Maire, Cat. PL Maroc. 3 767 



(1934). 

 Anacyclus valentinus subsp. dissimilis var. eudissimilis Maire in Jahandiez & Maire, Cat. PL Maroc. 3 : 



767 (1934), nom. illeg. 



Illustrations: Fig. 191, J. Schkuhr, 1808 : tab. 2546; Reichenbach, 1854: tab. 999, fig. 4; Cusin & 

 Ansberque, 1873 : tab. 148 : Coste 1903 : 348; Cadevall & Salient, 1917 : fig. 1423; Bonnier, 1922 : tab. 

 297, fig. 1475; Quezel & Santa, 1963 : tab. 975, fig. 2854. 



Capitula apparently discoid or subradiate, 12-0-18-0 mm in diameter. Outer florets ?, tubular to 

 ligulate, yellow, 3-5-8-0 mm long. Cypselas c. 4-0 x 3-4 mm, obovate with broad wings and usually 

 pointed auricles. 



Hybrid collections (Anacyclus x inconstans and A. x valentinus): 171 collections, mostly from north 

 Morocco, north Algeria and Spain. 



Distribution and ecology: See Fig. 23. Anacyclus homogamos and its hybrids, A. x inconstans and 

 A. x valentinus, are amongst the commonest roadside and field weed species of the western 

 Mediterranean region, particularly in Morocco. The normal habitat is disturbed ground, especially 

 in sandy and rocky places, particularly in montane regions of south-west Morocco but extending 

 into north Morocco, Algeria, Tunisia and Spain. 



It is not possible to demonstrate to any great extent, from herbarium material, the origin and 

 spread of hybrids except to report that they have a sympatric distribution with either or both of 

 the parental species. Along the eastern coast of Spain and the Mediterranean coast of France, 

 some populations consist almost entirely of A. x valentinus L. Mixed populations of A.x 

 valentinus, A. clavatus and A. discoideus occur around Oran in north-west Algeria, as is shown 

 particularly by the collection of Faure, and in the Moyen Atlas Mountains, as is shown in the 

 collections of Humphries, Jury, Mullin and Richardson. 



7. Anacyclus linearilobus Boiss. & Reuter 



Pugillus : 52 (1852). Orig. coll.: Reuter s.n., Alger: Oran, inter la Stidia et Mostaganem in arenosis 

 (G, holotypus). 

 Anacyclus acutilobus Durieu ex Boiss. & Reuter, Pugillus : 52 (1852). Orig. coll.: Balansa 689 (27 June 



1852), Alger: Sables a voisinant la Batterie espagnole, pres d'Oran (MPU, holotypus, BM, 



C, E, G, K, LE, P). 

 Anacyclus rubricantes Durieu ex Boiss. & Reuter, Pugillus : 52 (1852). 



Illustrations: Figs 2, 4, 24. Quezel & Santa, 1963 : 975, fig. 2853. 



