C. J. HUMPHRIES 



40 



km 500 



Fig. 23 Distribution of Anacyclus X valentinus () and A. x inconstans(^). 



Annual; stems 15-0-70-0 cm, erect, usually divaricately branching above, but occasionally from 

 the axil of a 'rosette' leaf, rather striate, deep red, glabrous below to lightly pubescent above. 

 Leaves 4-0-12-0 (-15-0) x 1-5-5-0 (-7-0) cm, ovate in outline, bi- to more rarely tripinnatisect, 

 usually sessile above to long petiolate below; primary lobes in 4-9 opposite to subopposite pairs, 

 with internodes up to 3 cm long; ultimate segments broad, acuminate, fleshy, rhachis narrow; 

 peduncle leaves (bracts) 2-4-0x0-5-1 -Ocm, pinnatisect, sessile, segments linear-acuminate. 

 Inflorescence a lax, corymbose cyme, usually with 2-4 capitula; peduncles erect, branching from 

 the middle of the stem, 7-23 cm long, distinctly clavate and hollow at the apex, pubescent 

 (especially when young) to sometimes glabrous at maturity. Involucre 8-0-17-0 mm in diameter; 

 involucral bracts in three rows, 4-5-7-0 x 1-5-2-5 mm, narrow-triangular to rectangular or ovate- 

 oblong, acute, somewhat membranous, but with a thick, pale green to brown centre vein and 

 narrow, scarious margins, invested with fairly long white hairs; receptacular scales 4-0-6-5 x 

 2-8-3-2 mm, obtrullate to narrowly obovate, cuspidate at the apex, with sharp spines at maturity, 

 canaliculate above, flat below, usually pale-brown, glabrous. Ray florets white, reflexed at 

 anthesis; ligules (5-0-) 8-0-1 6-0 x (3-0) 3-5-7-0 mm, acuminate to bifid, tube 1 -8-2-5 x 1-0-1-3 mm, 

 anterior-dorsally compressed, with broad ovate wings, female fertile. Disc florets 4-5-5-2 x 

 c. 1-0 mm, infundibiliform, lobes equal in the outer series, but two distinctly longer than the 

 rest in the inner series, wings broad, the basal part enlarged into an anterior circular appendage 

 up to c. 1 mm wide, extending over the cypsela. Styles 3-5-3-8 mm long, branches 0-5-0-8 mm 

 long. Stamens 3-5-4-0 mm long, anthers c. 1-8 mm long. Cypselas dimorphic, anterior dorsally 

 compressed but subrectangular with thin hyaline wings, those of the ray florets broadly ovate, 

 3-5-4-0 x 3-0-3-5 mm, the wings with rounded auricles 1-1-3 mm wide, those of the disc florets 



