REVISION OF ANACYCLUS 135 



Observations: This species is easily recognized by its characteristic broadly winged, scarious 

 cypselas. It can be distinguished from its sister-species, Anacyclus nigellifolius, by the truncate 

 base of the corolla tube, the broader cypsela wings and the broad rhachis of the upper leaves. 



Distribution and ecology: See Fig. 27. Anacyclus latealatus grows in fallow fields and steppic 

 communities around 1100m. It is known from only the south-west Turkish vilayet of Burdur. 

 According to Grierson (1975) the species is endemic and probably Irano-Turanian. 



Collections: Known only from the holotype. 



9. Anacyclus nigellifolius Boiss. 



Diagn. PL Or. I, 2 (11) : 13, t. 14 p. 267 (1849) ; Bois., Fl. Or. 3 : 322 (1875). Orig. coll.: Haussknecht s.n. 

 (April 1867), Mesopotamia, in gracuis, Dara (G, holotypus, BM, JE, Z). 

 Anacyclus nigellifolius subsp. orientalis Grierson in Notes R. hot. Gdn. Edinb. 33 : 41 1 (1975) ; Grierson 



in Davis, Fl. Turk. 5 : 223 (1975). Orig coll.: Sintenis 817, Turkey: Urfa, Nemrut Dagh (LD, 



holotypus, BM). 



Illustrations: Figs 2, 4, 26. Bouloumoy, 1930 : tab. 212, fig. 1 ; Davis, 1975 : 267, fig. 14, no. 13. 



Annual; stems erect, slender, 10-0-20-0 cm long, simple or sparsely branched from, or above the 

 middle, rarely much branched from below, glabrous. Leaves sessile, sparsely villous, bi- 

 pinnatisect to pinnatisect, 1 -5-3-0x0-3- 1-5 cm; primary lobes in 5-6 subopposite pairs; ultimate 

 segments 2-0-12-0x0-3-10 mm, acute; rhachis cuneate, thickened towards the base. Inflor- 

 escence monocephalic or a very loose corymbose cyme, peduncles usually emerging from or 

 above the middle of the stem, clavate below the capitulum at maturity, distinctly villous above. 

 Involucre 5-0-13-0 mm, turbinate-hemispherical; involucral bracts in three series, ovate acute 

 in outer series to obovate obtuse in inner series, 4-5-7-0 xc. 2 mm, villous, distinctly centrally 

 veined, green-brown towards apex, scarious, hyaline at the margin; receptacular scales obovate, 

 acuminate, 5-0-7-0 x 2-0-2-8 mm, slightly caniculate, scarious, but thickened slightly above, 

 glabrous. Ray florets white, ligule 4-5-6-0 x 1-0-3-0 mm, 2-3 fid; tube 3-5-4-8 x 1-0-1-2 mm, 

 anterior-dorsally compressed with narrow wings, persistent at maturity. Disc florets 3-5-4-0 mm, 

 base broadened into large circular appendage 2-5-3-0 mm wide covering top of cypsela on the 

 anterior side, lobes equal. Styles 3-5-3-8 mm long, style branches 0-5-0-8 mm long. Stamens 

 c. 4-0 mm, anthers 1-8-2-0 cm long. Cypselas monomorphic, somewhat rounded to obovate, 

 4-2-4-8 x 3-0-3-6 mm, pale brown; lateral wings extremely thin, transparent, scarious, 1-0- 

 2-0 mm wide, with rounded auricles. Pappus coroniform to virtually absent, contiguous with 

 wings. 



Flowering period: May-June. 



Observations: This species is similar in habit to Anacyclus latealatus and is its sister species, but it 

 has more erect stems, less elaborate leaves, and rounded auricles on the cypselas. It differs from 

 its congeners by the curious overlapping lobe at the base of the disc corollas, which is akin to that 

 found in the monotypic Leucocyclus formosus Boiss., in which the two overlapping lobes clasp the 

 top of the cypselas. 



Variation: Stem size, stem branching, size of involucre and florets are subject to considerable 

 variation. Grierson (1975ft) distinguishes two subspecies, nigellifolius and orientalis, which differ 

 in these characters, the generally more reduced form occurring in Hatay in southern Turkey. 

 This variation, however, is continuous and not distinctly geographical, since plants with all the 

 size-differences can be found in the Antilebanon, especially on Mount Hermon. The species is 

 polymorphic, with each population differing slightly in genetic constitution. I do not think that 

 this variation is worthy of taxonomic recognition. 



