REVISION OF ANACYCLUS \\\ 



11. Lower leaves 1-2-pinnatisect 



14. Disc corolla-tubes articulating normally with the ovary; cypselas with a lacerate 

 corona; rhachis of upper leaves broad 



15. Cypsela wing 2-5-2-75 mm wide; Turkey 8. latealatus (p. 134) 



15. Cypsela wing 0-3-0-6 mm wide; Algeria 7. linearilobus (p. 131) 



14. Disc corolla-tubes expanded into an orbicular flattened disc at the base of the dorsal 



margin, overlapping the ovary; cypselas ecoronate; rhachis of upper leaves narrow 



9. nigellifolius (p. 135) 



Sect. 1. PYRETHRARIA DC. 



Anacyclus sect. Pyrethraria DC., Prodr. 6:15 (1838). 



Anacyclus sect. Leucocyclus Batt. & Trabut, Fl. Alger. 3 : 453 (1890). 



Herbaceous perennials; stem reduced to a basal, woody subterranean caudex; leaves hetero- 

 morphic, tripinnatisect to bipinnatisect; radical leaves occurring in basal rosettes. Peduncles 

 many, prostrate, in annual rosettes. Ray florets white, with a red stripe below. 



1. Anacyclus pyrethrum (L.) Link 



Enum. Hort. Berol. Alt. 2 : 344 (1822); Cass., Diet. Sci. Nat. 34 : 102 (1825). 

 Anthemis pyrethrum L., Sp. PL : 895 (1753). Orig. coll.: Herb. Linnaeus 1016. 18. 'Pyrethrum 12' 



(LINN, lectotypus). 

 Anacyclus pseudopyrethrum Ascherson in Bonplandia 6: 123 (1858) Orig. coll.: Hohenacker, Arznei- 



u. Handlespflanzen 119 (holotypus destructus). 



Illustrations: Figs 2, 12. Edwards, 1799 : tab. 462; Schlectendal & Guimpal, 1833 : tab. 187; Meigen, 

 1842 : tab. 131e; Reichenbach, 1854 : tab. 999, fig. I, 1-3; Berg, 1861 : tab. 51, fig. 390; Artus, 1862-74 : 

 tab. 144; Argenta, 1864 : tab. 175; Ball, 1878 : tab. 24; Bentley & Trimen, 1880 : tab. 151 ; Hager, 1887 : 

 tab. 775; Pabst, 1888-90 : tab. 112; Quezel & Santa, 1963 : tab. 97, fig. 2849. 



Common names: Agargarha, Agonthas, Akurkurka, Aoud el Athas, Bertram, Guenthous, Igneus, 

 Manzanilla, Pellitory, nupeGpu, Piretro, Pyrethre, Pyrethri, Roman Pellitory, Spanish Chamomile, 

 Spanish Pellitory, Tagendest, Tigenthast. 



Perennial; stems fused with roots to form a submerged, woody caudex. Leaves heteromorphic, 

 slightly pubescent ; rosette leaves obovate in outline, petiolate, tripinnatisect, 3-0-14-0 x 0-5-3-0 cm 

 with tiny, acute, linear ultimate segments; primary lobes in 4-9 opposite or subopposite pairs; 

 rhachis slightly caniculate, cuneate at the base, the old veins often tough and persistent; peduncle 

 leaves sessile, or shortly petiolate, bi- to rarely tripinnatisect, 1-0-3-0x0-3-1 -5 cm, primary lobes 

 in 1-6 subopposite pairs. Inflorescence a contracted corymbose cyme with individual creeping 

 peduncles emerging from the centre of the caudex; each peduncle 6-30-0 cm long with 1-3 

 branches and terminal capitula, sparsely to densely villous, usually red, terete below the capitulum. 

 Involucre 7-0-22-0 mm in diameter; involucral bracts in three rows, narrowly triangular, vivid 

 green in the centre above, somewhat paler below with a distinct, but thin, dark-brown erose 

 margin; receptacular scales obcuneate, carinate to canaliculate, the apices broadly obtrullate to 

 cuspidate and overlapping the cypsela at maturity, somewhat membranous at the margin but 

 extremely tough at the centre and above and often with a distinct central vein. Ray florets white, 

 with a deep red stripe below; ligule 7-0-13-0 mm long, 3-0-4-5 mm wide, tube 3-0-5-0 mm long, 

 0-7-2-5 mm wide, anterior-dorsally compressed with narrow lateral wings, ^-fertile but occasion- 

 ally with vestigial anthers, articulating terminally on the ovary. Disc florets 3-0-5-0 mm long, 

 anterior-dorsally compressed, lobes narrow triangular, somewhat cucullate and equal in size. 

 Styles c. 3-5 mm long, the style arms 1-2-1-6 mm long. Stamens 3-9-4-1 mm long, anthers c. 2-0 mm 

 long. Cypselas obcuneate to broadly ovate, 3-0-4-0 x 2-8-4-0 mm, pale grey to dark brown, 

 sometimes purple, sphacelate; lateral wings thick, coriaceous, protruding apically with 2 

 acuminate auricles, 0-3-0-7 mm wide. Pappus absent. 



Flowering period: Mainly May to August. 



