118 



C. J. HUMPHRIES 



Distribution and ecology: See Fig. 15. Anacyclus monanthos extends from the Nile delta in north- 

 east Egypt to Ghardaia in the desert south of the Atlas Saharien in Algeria. Both subspecies are 

 pioneers of sandy soils. In this context it is interesting that subsp. cyrtolepidioides is the dominant 

 weed, replacing A. x valentinus, in the sandy wastelands of Biskra and Ghardaia in Algeria and 

 around Gabes and Gafsa in Tunisia. 



Fig. 15 Distribution of Anacyclus monanthos subsp. monanthos () and subsp. cyrtolepidioides (A). 



a. Subsp. monanthos 



Santolina terrestris Forskal, Fl. Aegypt.-Arab. 147 (1775). Orig. coll.: Forskdl (1762), Cent. 5, no. 72, 



Aegypto: Alexandria in vicinia columna Pompeyi (C, holotypus). 

 Anacyclus alexandrinus Willd., Sp. PI. 3 : 2173 (1800). Orig. coll.: Herb. Willdenow 16307 (B, lecto- 



typus). 



Anthemis arabica Viv., Fl. Lib. : 56 (1824), non. L. 

 Hiorthia alexandrina (Willd.) Less., Syn. Gen. Comp. : 258 (1832). 

 Cyrtolepis alexandrina (Willd.) DC., Prodr. 6:17 (1838). 

 Cyrtolepis alexandrina (var.) P glabra DC., Prodr. 6:18 (1838). 

 Cyrtolepis alexandrina DC. var. nov. Coss. in Bull. Soc. Bot. Fr. 36 : 95 (1889). 



Illustrations: Figs 2, 4, 14. Delile, 1814 : tab. 48; Gubb, 1913 : tab. 13; Quezel & Santa, 1963 : tab. 97, 



fig. 2850. 

 Common names: Sorret el Kebch. 



Peduncles prostrate, emerging from the axils of the rosette leaves or from the axils of receptacular 

 bracts of syncephalous capitula, frequently swollen and coalescing into groups of 2 or 3. Central 

 capitula present, occasionally 2-4 coalescing into a syncephalum or forming a congested inflores- 

 cence. 



Collections: 89 collections were examined, predominantly from the north desert areas of Libya 

 and Egypt. 



b. Subsp. cyrtolepidioides (Pomel) Humphries, stat. nov. 



Anacyclus cyrtolepidioides Pomel, Nouv. Mat. Fl. All. : 54 (1874). Orig. coll.: Kralik 248, Gabes (BM, 



E, G, LE, Z, isotypi). 



Anacyclus valentinus sensu Desf., Fl. All. 2 : 285 (1799) sphalm., non. L. 

 Cyrtolepis monanthos Less, in Linnaea 6 : 166 (1831). 

 Anacyclus mauritanicus Pomel, Nouv. Mat. Fl. All. : 55 (1874). Orig. coll.: Pomel s.n. Hautes steppes 



et montagnes des Ksous Mkraoula, El Beida, Aflou, Ksel (AL, holotypus; MPU, isotypus). 



