52 J. H. PRICE, I. TITTLEY & W. D. RICHARDSON 



Baixo Alentejo: 



Sines: Ginsburg-Ardre (1963); Ardre (1970); Ardre (1970), l-2/vii/1961, 4.X.1963. 



Vila Nova de Milfontes: Dizerbo (1954); Rodrigues (1963) and Ardre (1970) both citing 

 Welwitsch, s/n, iv.1848 (LISU P46430, P46433); Ardre (1970). 



Algarve: 



Sagres: Ginsburg-Ardre (1963); Ardre (1970); Ardre (1970), 27.ii.1960. 



Praia Da Rocha: Palminha, no data; Ardre (1970); Ardre (1970), 25.ii.1960, l.x.1963. 



Praia Do Carvoeiro: Palminha, no data; Ginsburg-Ardre (1963); Ardre (1970); Ardre 

 (1970), 15.viii.1960 and (J. Feldmann), 2.X.1963. 



Ria De Faro: Palminha, no data; Ardre (1970). 



Doubtful location: Torgulho: Herb. R. J. Shuttleworth, rocks, vi.1837 (BM). [Possibly a 

 location near Almada, south side of the Tejo (Tagus), north of Sesimbra; hence, Estremadura.] 



Portugal general: Rodrigues (1 963), aestival in Atlantic, iv to x; Ardre (1970, 1971), moderately 

 wave-beaten to sheltered shores, base of lower littoral and in infralittoral, on rocks covered by 

 sandy sediments, often in very well lit positions. 



Relatively well worked for some time, Portugal has recently been subjected to detailed long- 

 term study by Ardre (1970, 1971). Some Padina populations there are of considerable longevity, 

 e.g. that at Vila Nova de Milfontes, sampled by Welwitsch (specimens dated 1848) and Dizerbo 

 (1954), and noted by Rodrigues (1963) and Ardre (1970). There is indication that seasonal 

 periodicity is now beginning to show rather less consistency than further north; many more 

 collections of erect fronds date from earlier (February to April) and later (October) than is the case 

 for English and French populations. Ardre's (1971) information indicates no real difference in 

 habitat requirements between British and Portuguese populations. Both demand shelter from 

 direct strong wave action, although consistently wet conditions (usually standing water) are 

 necessary; both appear in the lower littoral and shallow infralittoral, on rocks covered with 

 detritus. 



Evenly spread geographically, the available data are not sufficiently extensive for certain 

 recognition of Padina distribution foci along the Portuguese coast. However, except for the single 

 record from Douro Litoral, all the available information derives from the southernmost coastal 

 provincias, Estremadura, Baixo Alentejo, and Algarve. Ardre's recent work was sufficiently 

 detailed and geographically widespread to demonstrate that this was not merely a matter of chance 

 distribution of collections, but had some real meaning in terms of Padina biogeography in Portugal. 

 Given tolerable local environment, Padina clearly grows better the further south it occurs along 

 Atlantic shores, towards the Mediterranean. The absence of records from the eastern half of 

 Algarve, beyond Faro, and from the whole of Huelva (see Atlantic southern Spain) is probably to 

 be explained by lack of adequate firm, detritus-covered substrata; both regions are very swampy 

 and sandy. 



Atlantic southern Spain: Huelva and Cadiz 



Huelva : 



No data traced; see note to Portugal. 



Cadiz: 



Chipiona: Seoane-Camba (1965), viii, on rocky platform, somewhat covered by sand and mud. 



La Caleta (Cadiz): Seoane-Camba (1965), vii, small specimens towards low water. 



Cadiz: Plantas de Andalucia, 1803, Leblech & Roxas Clemente, no. 313, 'Variedad de 312' 

 [young P. pavonica], no. 312 (MS Clemente) [older P. pavonica], on rocks by the castle of Sn. 

 Sebastian in Cadiz, mid-April, most abundant, Herb. R. J. Shuttleworth (BM); 161, most 

 abundant on rocks by the castle of San Sebastian in Cadiz, 29.iv (MS Clemente) (K in BM); 

 prope Gades, Dunze, 1803, Mertens, Roemer in Herb. R. J. Shuttleworth (BM); Clemente (1807) 

 (as Fucus pavonius L.); Seoane-Camba (1965), stn 79; specimens with no other data (two very 

 young, one older) (K in BM). 



