50 J. H. PRICE, I. TITTLEY & W. D. RICHARDSON 



bottoms, in small hollows between raised boulder arms, especially below Socoa fort; Hamel 

 (1939) (Bory); Evans (1957), widespread growths. 



Basse Pyrenees, general: 



D'Orbigny (1820), coasts of Gulf of Gascony (Basse Pyrenees), level two (1-5 m below MHW 

 to 4-5 m below MLW), common; Sauvageau (1897), fond du Golfe de Gascogne (and therefore 

 Basse Pyrenees), annual plant only once found in winter, as very small tuft among Cutleria, by 

 contrast, frequent in summer; Feldmann & Lami (1941), in sheltered stations, on bottoms covered 

 by mobile substrata, and in sunny, sandy-bottomed hollows, well-developed. 



There is a long history of original data on Padina from the rocky coasts of this most southerly 

 area of Atlantic France. The considerable ecological and phenological data, much greater than for 

 adjacent parts of France and Spain, are unequalled in quantity or detail south to the coasts of 

 Portugal and Atlantic southern Spain. The generalised location of the earliest records (D'Orbigny, 

 1820) to the Gulf of Gascony must be taken to include Basse Pyrenees; a precisely localised and 

 dated record for only 30 or so years later is known and there may be many other specimens in 

 French herbaria that would add to, but not materially alter, the picture. For example, there is 

 probably an approximately contemporary Bory specimen in Paris, since Hamel (1939) quotes 'St. 

 Jean-de-Luz (Bory)', signifying a specimen seen. Sauvageau's (1897) general statement for the Gulf 

 of Gascony (see Landes) also applies to Basse Pyrenees, being supported by Hamel's (1939) 

 quotations from Sauvageau herbarium specimens and by precise data published elsewhere by 

 Sauvageau (1920). Modern records (1940-70) indicate no significant recent change in the situation, 

 on this coastline at least. Ecological details in modern work indicate the general requirement for 

 standing wet conditions (best development in pools and channels); detrital conditions over rock; 

 and (generally) strong insolation. On these coasts, there is good general agreement with the usual 

 spring/summer/autumn sequence for the upright frond. Sauvageau's (1897) seems to be the only 

 attempt to follow events during winter and early spring in this area. 



Iberia 



Northern Spain: Guipuzcoa to Pontevedra 



Guipuzcoa : 



D'Orbigny (1820), Gulf of Gascony (and therefore Guipuzcoa), level two (1-5 m below 

 MHW to 4-5 m below MLW), common; Sauvageau (1897), fond du Golfe de Gascogne (and 

 therefore Guipuzcoa), annual plant only once found during winter, as very small tuft among 

 Cutleria, by contrast, frequent in summer. 



Santander: 



San Vicente de la Barquera: Sauvageau (1897); Seoane-Camba (1965). 

 Cabo Mayor: G. T. Boalch, 20.vii.1966. 



Oviedo: 



Gijon: Sauvageau (1897); Miranda (1931), area from outfall of La Nora, east of Cabo de San 

 Lorenzo, west to Cabo de Penas, Ria de Aviles, and Playa de Xago, frequent in pools, upper 

 littoral, fruiting in summer; Seoane-Camba (1965). 



Lugo: 



Rivadeo [ = Ribadeo] : Sauvageau (1897); Seoane-Camba (1965). 



La Coruna: 



Sauvageau (1897); Seoane-Camba (1965). 



Pontevedra: 



Bay of Vigo: Seoane-Camba (1958), amongst Enteromorpha ramulosa, in inlets. 



