THE DISTRIBUTION OF PADINA PAVONICA 37 



perhaps old wartime defences, reaches down virtually to low-tide level. Thickly covered by slimy 

 detritus, the boulders retain standing water at intervals; their surfaces are often well-shaded and 

 damp, even when above standing-water level. The boulder area, and shallow mud/shingle based 

 large pools at its inshore fringe, form a suitable environment for Padina, but none was detected 

 during careful examination in 1971 (J. H. P.; I. T.). Likely areas on and near Cap Gris Nez, and 

 elsewhere between Calais and Boulogne, were also unproductive. This lack is enigmatic; it could 

 be taken as a recession period, but that would not account for the similar absence of earlier 

 records, other than of drift plants, for the area. Strong wave-action and fetch may be a part- 

 explanation, since the English coast analogue of the Artois projection is the South Foreland which, 

 rather less directly affected by long fetch, still lacks a satisfactorily established history of early or 

 recent Padina records and carries no current populations (Price & Tittley, 1972). 



We have been unable to locate the 'Giard' source mentioned by both Debray and Chalon; 

 Giard was at this time Director of the Wimereux laboratory, so that specimen labelling or verbal 

 communication may have been involved. 



Somme 



No traceable records for the area. 



Since they are beyond the usual northern limits for consistent presence of Padina, the 

 continental areas so far dealt with (Pas de Calais; Nord; Somme; Netherlands; Belgium) constitute 

 the analogue of Essex, the Thames Estuary, Kent, and East Sussex. 



Seine Maritime 



St Jouin : 



Debray (1899), flat rocks, middle zone, based on Bernard (unpublished - see text); Chalon 

 (1905), Bernard; Coulon (19 12), pools, summer-autumn; Mail& Senay (1957), Bernard, '.. .Espece 

 fugace . . .'. 



Etretat: 



Wuitner (1921, 1946), in le Chaudron; Mail & Senay (1957), middle zone, au Chaudron, '. . . 

 Espece fugace . . .', based on Wuitner. 



Fecamp to Point e du Hoc, general: 



Mail & Senay (1957), rare, '. . . Espece fugace . . .', E. [bran], quoting from his unpublished 

 catalogue. 



Attached material has been recorded, but rarely; Seine Maritime and Pas-de-Calais have 

 much in common, since apparently ideal locations lacked and still lack plants. The area on the 

 north side of Fecamp Harbour (J. H. P. and I. T.; September 1972) is typical of these; flanked 

 by shingle, the location itself is of low boulders and flat slabs, the latter much eroded into declivities 

 and hollows, with standing water and muddy detritus. Taoniaatomaria was luxuriant and abundant, 

 but Padina was absent. Plants were also then lacking from Etretat area, although reported there 

 by Wuitner in 1 92 1 . Ebran and Bernard, the original sources of data quoted for this area, were local 

 botanists whose lists of marine algae were never published, although Ebran produced a catalogue 

 of vascular plants for the region of Le Havre and Bernard published a field excursion report which 

 referred generally to algae. Bernard was said to have inherited Ebran's algal list and incorporated 

 his own data with the intention of publication (Mail & Senay, 1957). 



Eure 



No precisely located records: 



Normandie* general: 



Lyle (1923); Chauvin in Suringar, in Herb. Weber-van Bosse, on rocks, all Normandy coasts 



* A name of variable application; we have taken Normandie to include Eure, Calvados, and Manche. Some 

 authorities also add Ille-et-Vilaine, which we have included in Brittany. 



