124 C. J. HUMPHRIES 



(1924) Orig. coll. : Braun-Blanquet & Maire s.n.(\92l), Grande He de Mogador. Paturages sablonneux 



entre Tiflet et Camp Monod (AL, holotypus). 



Anacyclus radiatus var. typicus Fiori, Nuov. Fl. Anal. Ital. 4 (2) : 650 (1927), nom. illegit. 

 Anacyclus radiatus var. typicus subvar. concolor Maire in Jahandiez & Maire, Cat. PI. Maroc. 3 : 766 



(1932), nom. wd. 



Illustrations: Figs 2, 4, 17. Miller, 1760 : tab. 73; Reichenbach, 1854 : tab. 999, fig. Ill, 12-14; Cesati, 

 Passer & Gibelli, 1867-86 : tab. 79, figs 5 b-e; Cusin & Ansberque, 1873 : tabs 147, 148; Regel 1882: 

 tab. 1074; Coste, 1903 : p. 348, fig. 1961 ; Fiori & Paoletti, 1904 : 426; Cadevall & Salient, 1917 : p. 278, 

 fig. 1422; Bonnier, 1922 : tab. 297, fig. 1474; Bouloumoy, 1930 : tab. 217, fig. 4; Post, 1933 : p. 58, 

 fig. 426; Negre, 1962 : p. 285, tab. 124; Zhangeri, 1976 : tab. 143, figs 5237, 5239, 5240; Haslam, Sell & 

 Wolseley, 1977 : tab. 45. 



Common names: El-Guentouss, Marguerite des Doukkala. 



Ligules yellow, or yellow with a red stripe below. Pappus 0-2-0-5 (-1-0) mm long on anterior, 

 (adaxial) face. 



Collections: 329 collections, predominantly from Spain, Morocco and France. 



b. Subsp. coronatus (Murb.) Humphries, stat. nov. 



Anacyclus radiatus var. coronatus Murb., Contr. Fl. Maroc 2 : 55 (1923). Orig. coll.: Ibrahim (20 May 



1889), Oued Tizi (LD, lectotypus, LE). 

 Anacyclus radiatus var. ochroleucus Ball in J.Linn. Soc. (Bot.) 16 : 504 (1878). Orig. coll.: Lowe s.n. in 



arenosis maritimus prope Mogador (K, holotypus). 

 Anacyclus exalatus Murb. in Bot. Notiser 1923: 61 (1923); Contr. Fl. Maroc 2: 55 (1923). Orig. coll.: 



Ibrahim s.n. (June, 1877), in herb. Cosson, prope oppidum Agadir imperii maroccani meridion alis 



(LD, lectotypus). 

 Anacyclus medians Murb. in Bot. Notiser 1923 : 60 (1923); Contr. Fl. Maroc 2: 53 (1923). Orig. coll.: 



Murbeck s.n. (23 April 1931), in herbosis ad Aguedal prope urbem Marrakech (LD, holotypus, 



MPU). 

 Anacyclus submedians Maire in Mem. Soc. Sci. nat. Phys. Maroc 15 : 39 (1927). Orig. coll: Maire s.n. 



(24 April 1925), Hab. in arvis argillaceo-humosis humidis planitei Gharb prope Sidi-Yaya, ubi 



martio et aprili floret (AL, holotypus, RAB, P, MPU). 

 Anacyclus ifniensis Caballero in Trab. Mus. nac. Cienc. nat. Madr. (Bot.) 28 : 24 (1935). Orig. coll.: 



Caballero s.n. (16 June 1934), en la plana de Ifni (M, holotypus, MPU). 

 Anacyclus ifniensis forma viridis Caballero, in Trab. Mus. nac. Cienc. nat. Madr. (Bot.) 28 : 25 (1935). 



Orig. coll.: Caballero s.n. (17 June, 1934), in umbrosis marginum flumine Ifni (M, holotypus). 

 Anacyclus radiatus var. coronatus Murb. subvar. discolor Maire in Bull. Soc. Hist. nat. Afr. N. 26 : 211 



(1935). Orig. coll.: Maire & Wilzeck s.n. (1 April 1934), sables maritimes pres du Cap Ghir (AL, 



holotypus, RAB, P). 



Illustrations: Murbeck, 1923 : p. 54, figs a-h. 



Ligules white, or rarely white with a red stripe below. Pappus (0-3-) 0-5-1-2 mm long on 

 anterior (adaxial) face. 



Collections: 51 collections, particularly from the Sous valley in south-west Morocco and Lanz- 

 arote in the Canary Islands. 



5. Anacyclus clavatus (Desf.) Pers. 



Syn.Pl.2 :465 (1807). 



Anthemis clavata Desf., Fl. Atl. 2 : 287 (1799). Orig. coll.: Herb. Desf. (P, holotypus). 

 Anthemis tomentosa sensu Gouan, Obs. Bot. : 70 (1773), pro parte quoad holotypus autem non L. 



( = Anthemis). 



Chamaemelum tomentosum sensu All., Fl. Pedem. 1 : 184 (1785). 

 Anthemis pedunculata Desf., Fl. Atl. 2 : 288 (1799). Orig. coll.: Herb. Desf. (P, holotypus). 



