U-2 FOSSIL INSISTS OF THE BRITISH COAL MEASURES. 



the presenco of a wide interval between the former and the radius, and the wide 

 area occupied by the latter. 



The division of the radius into two unequal branches, with its symmetrical 

 double bifurcation, is also, I venture to think, a feature of primary importance. 



In the wide divergence of the radius and median, this species agrees with 

 Scudder's genus Spiloblattina ; but in this case the veins do not converge again to 

 enclose an elongated or oval area. 



Archimylacris, sp. indet. Plate VII, fig. ">. 



1911. Archimylacrie, sp inilct., UolUni. (^duirt. Journ. Geol. Six-., vol. Ixvii, p. !<>:>, pi. x, HL;. :l 



TI/JH_>. A fragmentary wing, lacking the apex and the base ; Museum of 

 Practical Geology, .lermyn Street (no. 24o03). 



llni'i:;on a iid Locality. Base of Upper Coal Measures (Grellideg Level of the 

 Mynyddislwyn Vein) ; near Maes-y-cwmmer, Monmouthshire. 



Description. The specimen is much too fragmentary for any attempt at specific 

 determination. A portion of the distal margin is present, with three branches of 

 the subcosta. These are succeeded by eight straight veins, two at least forking, 

 which belong to the radius and median series. These are followed by an equal 

 number of veins partially hidden by the surface of the integument, which is much 

 wrinkled over this area. These, I assume, are parts of the cnbitus vein. The 

 wrinkling of the integument over the cubital area is very marked. 



The special interest of this wing-fragment is in its association with portions of 

 a leaf of ' 'orilniles. Scudder and others have commented on the general association 

 of the wings of Blattoids with leaves of (_'t-<lititi'x. but have not. so far as 1 am 

 aware, drawn attention to a feature which is well shown by this leaf, namelv, pits 

 on its surface. I have repeatedly found such depressions on the leaves of ('oriliiitt'* 

 in the Lancashire Coalfield, and in many cases seen the hollows occupied by the 

 shells of S/iii'in-l'ix /iKxillitx. The shallow pits on the Cordaites-leai associated with 

 this fragmentary wing show faint traces of a spiral, similar to the impression 

 of ,^/iii-ni'liix, and such shells were. 1 think, once attached to it. While the 

 Carboniferous Pdattoids may have Keen wholly phytophagous, we are led by our 

 knowledge of t lie living Blattids, especially by t lie common cockroach ( I'lTi/il'iiit'l'i 

 orientalis), to assume the contrary, and to regard them as more likely to have been 

 omnivorous, in which case the association of Blattoid remains with the leaves of 

 I 'm'lliiili'x bearing the sedentary Spvrovbis is easily understood. The association 

 :ilso lends support to the beliel that the Blattoids were semi-aqiiat ic in habit, or 

 lived in marshes and >wamps m which dccaved veivel at ion Formed a home for 

 Spii'orbis. 



