with Remarks on their Affinities. 58 



in GakuVy and six in Francoa, and alternating in both ge- 

 nera with the segments of the calyx. Both genera likewise 

 correspond in having barren filaments alternating with the fer- 

 tile ones : but in Galax they are united together in a tube, and 

 the anthers consist of a single cell, opening by a horizontal su- 

 ture. The capsule of both is composed of several loculaments, 

 with septiferous valves, and opening in the middle by means of 

 a longitudinal fissure. The cells unite at the centre, which is 

 destitute of a columella, the seeds, which are numerous and mi- 

 nute, being attached to the inner angle of each cell. The cells 

 are four in Francoa, and most commonly three, although some- 

 times also four in Galax. The stigma, which in both is really 

 compound, in the former is 4-lobed, with a perfectly distinct 

 base, while in the latter it is undivided, with connate lobes, their 

 presence being indicated by the furrows, which correspond in 

 number with the cells of the ovarium. The texture of the 

 leaves, their nervation, and the presence of glands terminating 

 the teeth, the leafless stalks, the spiked inflorescence, with single- 

 flowered pedicels, each being furnished at the base with a small 

 persistent bractca, are additional evidence of the intimate affi- 

 nity existing between these two genera. This small family, 

 which I have named Galacince, may be characterized as follows : 



GALACINvE. 



Cali/x 4-5-partitus, persistens. Petala 4-5, hypogyna, calycinis segmentis al- 

 terna, caduca. Stamina 10 v. 16, hypogyna, in tubuni apice 10-dentatum 

 connata, aut distincta : filamentis dentilmsve altemis lantum antheriferis. 

 Ovarium (e tribus v. quatuor tblliculis compositum) 3-4-loculare : ovulis in- 

 definitis. Stigma (e 3-4, compositum) iudivisum v. 4-lobum. Capsula 3-4- 

 locularis, 3-4-valvis : valvis medio septiferis. Placenta centralis nulla. 

 Semina numerosa, minuta, scobiformia, angulo interiori loculorum inserta ; 

 testa exterior laxa, membranacea, cellularis ; interim' tenuissima, albumini 

 arete adhaerens : albumen copiosum, carnosum. Embryo erectus, teres : co- 

 tyledones brevissimae : radicula longa, cylindracea, centripeta. 



Herbae (Americana?) perennes. Folia radicalia, simplicia v. lyiata^ pinnatijida-, 

 serrata : dentibus glandula coronatis. Flores terminales, copiosi, spicato-race- 

 mosi. Pedicelli M/tt/fori, basi bracteola persistente suffxdti. 



Obs — Filamenta in Galace 10, in tubum apice 10-dentatum connata: den- 

 tibus altemis brevissimis rotundatis crassiusculis antheriferis ; caeteris petalis 

 oppositis, elongatis, spathulatis, nudis : antherae infra apicem laterique inte- 

 rior! filamentorum insertae, uniloculares ! loculo transverse, rima bivalvi ho. 

 rizontali hiantes : stigma (e tribus arcti connatis) trigonum, trisulcum : cap- 

 sula columella destituta : semina utrinque mucronata, omnin* ^cw/wvr, De 

 Cand. Familia prope Philadelpheas et Saxifrageas locanda. 



