OF WASHINGTON, VOLUME XIV, 1912. 163 



SIX NEW GENERA OF NEARCTIC MUSCOIDEA. 



BY CHARLES H. T. TOWNSEND. 



The following six new genera are erected for as many de- 

 scribed North American species, which can not be placed in 

 any previously erected genera known to me. The reproductive 

 habits of four of these genera are known, and these four are 

 treated in a forthcoming paper on the female reproductive 

 system, eggs, and early stages of muscoid flies. 



Neophorocera, gen. nov. 



Erected for Phorocera edivardsii \V ill. (Euphorocera claripennis Coq. 

 pt.). The male has the second antennal joint hardly at all elongate, 

 while the female had it noticeably so. The length of the second antennal 

 joint thus becomes a secondary sexual character in this genus, which is 

 the first case of the kind known to me in this immediate group. In the 

 salmaciine and belvosiine flies the same secondary sexual character of 

 the second antennal joint is much more pronounced. 



Reproductive habit, oviposition of flattened macrotype egg on host. 

 The egg-chorion is honeycomb-reticulate and without operculum. The 

 uterus is absent, but the uterovagina of one of the dissected specimens 

 (TD394) showed an egg containing a fairly developed maggot. This 

 development of the maggot in the uterovagina is probably exceptional, 

 due to undue retention of egg from failure to find a suitable host. (TD394, 

 coll. by D H. demons, Aug. 22, 190S, Melrose Highlands, Massachusetts, 

 and determined by W. R. Thompson as Euphorocera claripennis Coq.). 



Type: Phorocera ed-wardsii Will., Scudder's Butterflies of 

 New England, vol. in, p. 1921. 



The genera Exorista, Tricholyga, Euphorocera, andC/ueto- 

 tachina have the egg-chorion without reticulation and fur- 

 nished with an operculum at the cephalic end. \Vinthciuia 

 appears to lack both reticulation and operculum. C/ueto- 

 lyga and Nemorilla appear to have the chorion reticulate and 

 without operculum, but are at once distinguished from Neo- 

 phorocera by their ciliate hind tibiae. 



Euacemyia, gen. nov. 



Erected for Acetnyia tibialis Coq. The frontal bristles stop at base of 

 antennae, the vibrissa- are inserted well above oral margin, the third an- 

 tennal joint of female is only about one and one-half times as long as second, 

 the cheeks of female are only one-fifth of eye-height, the apical cell is open 

 and ends well before wing-tip. In Acemyia the frontal bristles descend 

 nearly or quite to base of third antennal joint, the vibrissae are inserted 

 practically on the oral margin, the third antennal joint of female is fully 

 or more than twice as long as second, the cheeks of female are over one- 



