134 PROC. ENT. SOC. WASH., VOL. 21, NO. 6, JUNE, 1919 



fly, it is a comparatively simple matter to determine the corre- 

 spondence of the genital forceps of a primitive sawfly such as 

 Megaxyela (Fig. 27, "gb" and "eg") with the forceps called 

 gonopods (Crampton, I9i8b) in such primitive Mecoptera as 

 Merope (Fig. 35, "gb" and "eg"). In some Mecoptera as in 

 the one shown in Fig. 31, the basal segments of the genital for- 

 ceps "gb" unite to form a basal region bearing the distal segments 

 "eg" of the gonopods, or forceps, and in the same way, in some 

 sawflies, as in the one shown in Fig. 9, the basal segments of the 

 genital forceps "gb" unite to form a basal region bearing the distal 

 segments "eg" of the gonopods or forceps. This interpretation 

 of the parts seems so self-evident, that it is difficult to understand 

 how Newell, 1918, comes to such a different conclusion as to the 

 homologies of the parts, unless the wrong labels were attached 

 to her figures, and her tables of sclerites and appendages were 

 inadvertently placed under the names of the wrong insects in 

 some cases. Thus in her figure of the genitalia of Dolerus, Newell 

 would interpret the distal segment of the forceps "eg" of Fig. 39 

 (of the present paper) as "appendage IV" homologous with the 

 cerci of lower insects, although she correctly figures the cerci 

 of a male Dolerus elsewhere. The median plates with their 

 processes "pa" of Fig. 39, Newell would interpret as the homo- 

 logues of the distal segments of the gonopods "eg" (Fig. 31) of 

 the Mecoptera, while the basal region of the gonopods of the 

 Mecoptera ("gb" of Fig. 31) are homologized with the basal 

 ring "gg" of Dolerus (Fig. 39) by Newell, who regards the basal 

 ring "gg" of Fig. 39 as the sternum of the ninth abdominal seg- 

 ment in Dolerus, although it probably belongs to the tenth seg- 

 ment instead (the true ninth sternite being the large plate "ha" 

 of all figures), and it comes to have a ventral position in Dolerus 

 only secondarily, through a revolution of the copulatory appara- 

 tus about its long axis. The homologies proposed by Newell 

 for the Neuroptera, and other forms, are also not in accord with 

 the interpreptation of the parts given in a paper dealing with the 

 gonopods of these insects, Mecoptera, etc. (Crampton, 19185), 

 but it is not necessary to take up the discussion of the gonopods 

 of these insects here. 



Berlese, 1909, interprets the basal segments of the forceps of 

 Cimbex (Figs, i and 14, "gb") as the sternite of the tenth ab- 

 dominal segment, although they are clearly the homologues of 

 the basal segments of the gonopods of lower sawflies (Fig. 27, 

 "gb"), Mecoptera (Figs. 35,31, "gb"), etc. The distal segments 

 of the forceps (Figs, i and 14, "eg"), Berlese calls "stili" in Cimbex, 

 and applies the same designation to the styli of ephemerids 



