OF WASHINGTON, VOLUME XIII, 1911. 167 



Cynomyia, type morttioruni, long classed with the Sar- 

 cophagidae and even continued so to the present day by many 

 authors, was shown by Portchinski to belong unmistakably 

 to the calliphorine flies. Its maggots are almost identical 

 with those of Calliphora, it is oviparous, and evidently lacks 

 uterus. It thus goes in the subfamily Calliphorinse. 



The reproductive habits in the important subfamily Mesem- 

 brininae are well set forth by Portchinski. Mesembrina, type 

 mystacea, deposits on dung not over two dozen large eggs, 

 which are 4 mm. in length. These eggs, which look more 

 like small pupae, hatch in about twenty-four hours into mag- 

 gots with anal stigmata characteristic of the first stage. 

 These, Portchinski states, shortly change directly to the third 

 stage, entirely omitting the second stage, the stages being 

 identified on the characters of the anal stigmata. The maggot 

 period is very short. 



Metamesembrina, type meridiana, islarviparous, depositing 

 large maggots in dung. It was investigated in the Crimea. 

 A female was found to contain a large maggot 3 mm. long, 

 and alongside of it an egg of the same size. It was not fol- 

 lowed further apparently, but probably has the same habit as 

 Dasyphora next to be considered. 



Dasyphora, type pratorum, was found to have a remark- 

 able style of reproduction, then for the first time made known 

 outside of the Pupipara, and thus announced before this 

 habit was known to exist in Glossina. One large egg at a 

 time is retained in the uterus, and not only is it developed to 

 the maggot, but the latter is carried through its first and 

 second stages in the uterus and deposited in its third stage on 

 dung. It feeds to some extent before pupating, thus differing 

 from the deposited maggot of Glossina, which is said to feed 

 not at all outside. 



Here then are several types of reproduction in the subfamily 

 Mesembrininae, and the most specialized showing a remark- 

 ably close approach to that of the tsetse-flies, Glossina and 

 perhaps Glossinella, which I have considered a group of the 

 subfamily Muscinae, but which may yet prove to be more 

 closely related with the Mesembrininae. At all events it is 

 well established by Portchinski's work that the mesembrinine 

 flies, unlike the calliphorine, possess a true or incubating 

 uterus. 



Remarkable indeed are the results secured by Portchinski 

 in his investigation of Musca corrina, which show beyond 

 doubt that this species is not congeneric with Mtisca donies- 

 tica. It becomes necessary to found a new genus for its re- 



