HARRIS AND KUCIIS : OBSERVATIONS ON POLLINATION. 17 



corolla, varying sometimes to a greenish yellow. The large 

 anther, however, is quite different ; the proximal half being of 

 a greenish yellow, while the distal half has a more or less pro- 

 nounced purple color. Professor Todd, in his paper, does not 

 speak of the color of the anthers, but Fritz Mueller, in writing 

 of S. rostratum, says: "All the anthers, as I am informed by 

 Professor Todd, are of the same dull yellow color." All the 

 material examined by the writers from this locality shows a 

 decidedly different color for the distal half of the large stamen. 

 It seems hardly probable that material growing in Iowa should 

 show such a marked difference, but in case this statement is 

 not the result of an oversight on the part of Professor Todd, it 

 is of considerable interest. The anthers dehisce by terminal 

 pores, as is common in the genus to which the plant belongs. 



The two lower lobes of the corolla are produced into short 

 wings, which in the bud enfold the pistil and the large stamen, 

 which is clearly differentiated as such in the youngest buds in 

 which the stamens may be discerned by careful dissection. In 

 the bud the pistil lies immediately above the large stamen, but 

 upon the opening of the flower extends between the filaments 

 of the large stamen and that of the small stamen either to the 

 right or to the left. 



Professor Todd's statement is: "The pistil in any flower 

 turns toward the axis of the raceme." While in a general way 

 this is true, the statement might be more clearly expressed, 

 since it is only in the general direction of the pistil as a whole 

 that it points toward the axis of the raceme. 



The style is not inserted perfectly perpendicularly upon the 

 top of the ovary, but bends slightly downward from the longi- 

 tudinal axis of the flower. Professor Todd has overlooked this 

 point in his figure. Throughout the remainder of its course 

 until near the tip it is almost straight. Thus it will be seen 

 that the large stamen and the pistil are placed almost opposite 

 each other on the lower side of the flower. The angle between 

 their incurved ends, which approach within about three mm. of 

 each other, is about seventy degrees, thus causing them to point 

 toward opposite sides of the flower. Thus it will be seen that, 

 since the flowers are arranged alternately on the opposite sides 



5. Mueller, Fritz : Two Kinds of Stamens with Different Functions in the same Flower, Na- 

 ture, vol. XXVII, pp. 364, 365, 1883. 



2 -Kan. Univ. Sci. Bull., Vol. I. 



