WILLIAMS: MUD CRAB. PARTITION INTO SIX SPECIES 



SPECIES ACCOUNTS 



Panopeus austrobesus new species 



Figure 1 



Panopaeus herbstii. —Heller 1868:16 (Brazil, local- 

 ity). 



Panopaeus herbstii. — Ortmann 1893b:475 (part). 



Panopeus herbstii forma obesa. — Rathbun 1930:336 

 (part, specimens from south of Cabo Frio, 

 Brazil). 



Panopeus herbstii forma crassa. — Rathbun 1930:336 

 (part, specimens from south of Cabo Frio, 

 Brazil). 



Panopeus herbstii. — Coelho and Ramos 1972:190 

 (part).— Williams 1984:412 (part). 



Material examined. — Specimen lots in USNM re- 

 corded by Rathbun (1930) under P. herbstii (cata- 

 log number only) plus material added since that 

 time. 



Brazil: 47847, 47851, 59457, 59458, 59459, 

 59460,59461,59462,59465.-75661. 2M juv.; Ilha 

 Paqueta; W. L. Schmitt, 23 August 1925.— 

 75663. IF (ovig.); Ilha Govenador; W. L. Schmitt, 1 

 September 1925. — 75603. 5 juv.; Santos, on muddy 

 beaches between canals 4 and 5; W. L. Schmitt, 13 

 September 1925.— 75654. 3M, IF, 3 juv.; Santos 

 estuary; W. L. Schmitt, 13 September 1925. — 

 75653. 1M juv.; Paranagua; W. L. Schmitt, 2 

 October 1925. — 75662. 1M juv.; Paranagua; W. L. 

 Schmitt, 3 October 1925. 



Types. — The following are designated and labelled as 

 types deposited in the crustacean collection of the 



USNM: 59462. Holotype male, cl 26.5 mm, cw 38.7 

 mm, Paranagua, Brazil, sandy mud flats, under scat- 

 tared rocks; W. L. Schmitt, 3 October 1925. — 

 191147. 7M, 2F, same lot of specimens as above and 

 considered as paratypes. 



Diagnosis. — Carapace with few transverse lines of 

 granules, slightly coarsened granules on ocular and 

 hepatic regions and along anterolateral slopes 

 (length 64.2-72.9% width, x 68.8). -Coalesced first 

 and second anterolateral teeth of carapace separated 

 by very shallow notch, broadly rounded second tooth 

 usually but not always exceeded by acute first tooth; 

 remaining teeth outstanding in large adults, less so in 

 smaller specimens, third tooth with rounded to rec- 

 tilinear tip, fourth swept forward and acute, fifth 

 much smaller, acute and directed anterolaterad; arc 

 drawn along tips of first 4 teeth diverging laterally 

 from arc drawn along bases of notches between teeth 

 Chelipeds microscopically granular, sometimes more 

 conspicuously so along anterior edge of carpus, gran- 

 ules on outer surface of palm often arranged in 

 obsolescent rugose pattern. Major chela with teeth 

 on fixed finger varied in size, one or more distal to 

 level of basal tooth on dactyl enlarged, with cusps 

 raised above straight line drawn between angle at 

 juncture of finger with anterior margin of palm and 

 tip of finger (= length of fixed finger) (Fig. 2) ; cusps of 

 teeth on fixed finger tending to be well separated, 

 seldom worn, their external faces sometimes promi- 

 nent but not noticeably bowed outward from axis 

 of finger. 



Measurements in mm. — A set of measurements for 

 length and width of the carapace of this species is 

 summarized in Tables 1 and 2 and graphically rep- 

 resented in Figure 9. 



FIGURE I.— Panopeus austrobesus.— Holotype male from 

 Paranagua, Brazil (USNM 59462) a, Carapace in dorsal 

 view, b. Major chela in frontal view; 10 mm indicated. 



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