FISHERY BULLETIN: VOL. 81, NO. 4 



oyster bed 0.5 mi S of light at low tide; J. P. E. 

 Morrison, 6 July 1935.— 191149. IF (ovig.); 

 Wachapreague; 24 July 1943.— 63322. IF; Cape 

 Charles, near wharf; H. G. Richards.— 63323. IF 

 Guv.); Cape Charles; H. G. Richards.— 63324. 2F; 

 Cape Charles, near wharf; H. G. Richards. — 

 74600. 5M; York River, 6 ft; J. B. Engle, U.S. Bur. 

 Fish., 21 October 1936.— 75628. IF; Yorktown; J. 

 B. Engle, U.S. Bur. Fish— 75668. IF; Yorktown; J. 

 B. Engle, U.S. Bur. Fish. 



North Carolina: 62538,62539,62540.-60607. 1M; 

 Hatteras; A. J. Poole and R. Kellogg, May 1927- 

 191172. 2 M; Pivers I., Beaufort; J. B. Sullivan, 10 

 July 1978.— 84428. 2M; Fort Macon, Beaufort, 

 under stones in muddy zone just above low water on 

 breakwater; T. A. and A. Stephenson, Stn. 

 BM(N)39, 1947. 



South Carolina: 15726, 15741, 15762, 15780, 

 15782, 26150, 42881, 45502, 51022, 56265, 

 59938.-75667. 1M; Dewees I.; G. R. Lunz, Jr., 15 

 May 1934. 



Florida: 56838.— 99909. 2F; Bridge over Matan- 

 zas River, Crescent Beach; D. K. Caldwell et al, 1 

 December 1954.— 170017. 1M; Sebastian Inlet, 

 Indian River Co.; on sabellariid reef S side under 

 bridge, intertidal; R. H. Gore, 7 May 1974. 



Locality uncertain: 56840. 2 F; east coast of North 

 America; Boston Society of Natural History. 

 -75596. 1M, 5F (1 ovig.); O. Bryant, Boston Soc. 

 Nat. Hist. 



Diagnosis. — Coalesced first and second antero- 

 lateral teeth of carapace separated by shallow round- 

 ed notch, second tooth broader than but not so 

 prominent as first tooth; third and fourth antero- 

 lateral teeth curved anteriorly; fifth tooth much 

 smaller than fourth, acute and swept forward or 

 hooked anteriorly. Carapace length 66.8-77.0% 

 width, x 73.2. Chelipeds superficially smooth but 

 actually finely granulate on upper surface of carpus; 

 lower outer surface of palms light colored (pink, buff, 

 or yellowish, rarely mottled reddish); major chela 

 with cusps of teeth on fixed finger usually either 

 reaching or falling below straight line drawn between 

 angle at juncture of finger with anterior margin of 

 palm and tip of finger (= length of fixed finger); 

 anterior margin of palm with distance between crest 

 at base of dactyl and tip of condylar tooth lateral to 

 base of dactyl 0.70 or less length of fixed finger. 



Measurements in mm. — A set of measurements for 

 length and width of the carapace of this species is 

 summarized in Tables 1 and 2 and graphically rep- 

 resented in Figure 9. 



Color. — Dull olive, brown- green, dirty gray or slate 

 color, sutures between regions sometimes lighter, 

 and some tendency to variegated dark pattern on 

 lighter background. Chelipeds darker with variega- 

 tion coarse on upper parts, sometimes spotted claret 

 brown, blotching extending about halfway down 

 sides of articles, lower half and underparts light; 

 fingers black. Third maxillipeds of males and about 

 half of females with red or burgundy spot near base of 

 inner surface of ischium. (Specimens from Beaufort, 

 N.C.; Hay and Shore 1918; Rathbun 1930.) 



T]ype- locality. — "Inhabits oyster beds, &c . . . often 

 found on oysters (O. uirginica) in our markets" [by 

 implication the eastern United States] (Say 

 1817:58). Holthuis's (1979) selection of the 

 specimen figured by Say (1817, pL 4, fig. 3) asthelec- 

 totype for P. herbstii restricts the nominal species to 

 the common mud crab occurring on oyster bars of the 

 eastern United States. 



Known range. — The known geographic range of this 

 species, represented by the material listed above, is 

 shallow intertidal and subtidal waters of the eastern 

 United States from Boston Harbor, Mass., to Indian 

 River County, southeastern Florida. 



Remarks. — See general discussion below. 



Panopeus lacustris Desbonne 

 Figure 4 



Panopeus lacustris Desbonne 1867:28. 



Panopeus herbstii. — Gibbes 1850b:176 (part, Key 

 West material).— Smith 1869a:276 (part, the 

 Bahamian material). — Kingsley 1880:393 (part). — 

 Rathbun 1884:772 (part).— Benedict and Rathbun 

 1891:358 (part, the Bermuda, Caribbean, and 

 Brazilian material).— Young 1900:132.— Rathbun 

 1924:14.— Chace 1940:34.— Chace and Hobbs 

 1969:154, figs. 46c, 47. — Holthuis 1959:207.— 

 Edmondson 1962:277, fig. 20c (?).— Coelho 

 1966:163 (habitat).— 1970:53 (habitat).— Coelho 

 and Ramos 1972:190 (part).— Fausto Filho 

 1966:33 (occurrence). — Bonnelly de Calventi 

 1974:27 (list).— Markham and McDermott 1981: 

 1273 (list).— Williams 1984:412 (part). 



Panopeus Herbstii. — Smith 1869b:34 (part, list- 

 ing).— von Martens 1872:89.— Verrill et al. 

 1874:547 (part).— A. Milne Edwards 1880:13 

 (Brazil locality). 



Panopeus herbstii var. obesus Smith 1869 a: 

 278 (part, the Aspinwall material). 



868 



