FOLSOM: MOUTH-PARTS OF ANURIDA MARITIMA. 123 
Tay. IL. Figure 2, 3, 6,8; v. Stummer-Traunfels, ’91, Taf. I. Figuren 
4, 10) has a two-lobed galea and a four-lobed lacinia. 
In Japyx, thanks to Meinert’s figure (65, Taf. XIV. Figur 8), the 
muscles may be clearly homologized with those I (99, Plate 3, Figures 
20, 21) have described for Orchesella. As Meinert did not designate 
the muscles, I can simply say that they severally correspond with those 
labelled by me 3. add., 4. add., 10. add., 7. add. or 9. pr’t. add., and one 
muscle with both 4. pr’t. add, and 6. prt. add., while one of two others 
probably represents 8. ret. add. 
In Lepisma, according to Heymons (’97°, p. 592, Taf. XXX. Figuren 
13, 15, 17, 20), the fundament of the first maxilla forms the pal- 
pus, at the base of which appears a mesal lobe, which itself divides 
to form galea and lacinia. This account is, then, at variance with mine 
on Anurida, that of Uzel on Campodea, and that of Ayers for the Orthop- 
teran genus (Ecanthus, and is, so far as I know, unsupported by the re- 
sults of other authors. In fact, Figure 13 of Heymons even suggests 
that the palpus is a lateral lobe of the primary fundament, as I have 
found it to be in Anurida. As to the origin of the three first maxillary 
lobes, Uzel, Heymons, and myself disagree, as I have said. Uzel’s 
account agrees with mine, in so far as he makes the palpus a lateral 
evagination of the primary, or stipal, fundament ; and Heymons, like 
myself, derives both lacinia and galea from the inner lobe of a biramous 
appendage. 
In its final form, the first maxilla of Lepisma is easily recognized as 
homologous with that of other Thysanura, but approaches remarkably 
the same organ in Orthoptera, especially that of the Blattide. As in 
other Apterygota, the stipes (v. Stummer-Traunfels, ’91, Taf. II. Figur 
11; Muhr, ’77, Taf. VII. Figur 45) has a basal opening, cardo, distinct 
head, galea, and lacinia, and the origin of the muscles (Oudemans, ’88, 
p. 187) “findet man auch hier an Chitinstiicken im Kopfe.” The 
palpus in Lepisma, however, is five-jointed, as in Orthoptera. What I 
call galea and lacinia are also, in this particular case, named “ Aussen- 
lade” and “ Innenlade” by v. Stummer-Traunfels. 
Machilis is nearer than Lepisma to Campodea and Collembola in the 
structure of the first maxille. As may be seen from the figures by 
Oudemans (’88, Taf. II. Figur 27) and v. Stummer-Traunfels (’91, 
Taf. II. Figuren 8, 9, 10), the positions of the cardo, stipes, galea, 
lacinia, and palpus are exactly comparable in the three groups. The 
palpi in Machilis, to be sure, are seven-jointed, and a palpiger is present, 
as in Orthoptera, The structure identified by v. Stummer-Traunfels as 
