130 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 
It is not clear, then, whether the galea develops from the outer or 
the inner lobe of a biramous appendage, although Uzel’s account is, at 
least, not inconsistent with his description of the first maxille, which I 
have already criticised. Although Uzel does not state as much, his 
figures indicate the palpus to be an appendage of the primary funda- 
ment, as it is in Anurida. In this genus, however, no third branch ap- 
pears, as I have said; but, from analogy with the first maxilla, the inner 
of the two branches represents undifferentiated galea and lacinia. 
The rotation in a frontal plane of the second maxillary fandament of 
Campodea — which does not occur in Anurida— enables me to homo- 
logize the finished labium of Campodea with the apparently different 
labium of all other insects. If Uzel’s figures are compared with Figure 
12 of y. Stummer-Traunfels (91), it is easy to see that the embryonic 
structures by Uzel designated lim, (lacinia), Jem, (galea) and pmax, 
(palpus) are with hardly a doubt respectively represented in the adult by 
the parts which v. Stummer-Traunfels termed up. (“untere Mundplatte”), 
pl. (‘‘tasterformige Papille”) and pp. (‘‘ Tastwarze”’). These homo- 
logies, however, could never have been settled upon merely anatomical 
grounds. 
What Grassi (86°, Tav. IV. Figura 3), then, considered to be the 
under lip (/a. a.) of Campodea is but the anterior part of the true 
labium; the “labial palpi” (pa. lz.) are really galee borne upon a 
region representing the mentum, and the “labial papille” (pa. da.) 
are but modified palpi. As in Collembola, the labium is anteriorly and 
deeply cleft. 
Japyx is so close to Campodea that the same conclusions may doubt- 
less be applied to both genera. In Japyx the labium, as in Collembola, 
is split and bears a median sulcus (Grassi, 86°, Tav. III. Figura 21) 
much like that of Orchesella (Folsom, ’99, Plate 4, Figure 29). Ex- 
amining Figure 1 of v. Stummer-Traunfels (’91, Taf. II.), the lacinia and 
galea are clearly represented, as in Campodea; the true palpus, however, is 
but obscurely differentiated in the region behind the so-called palpus (pl.) 
and nearer the median plane. The eversible papillee of the anterior part 
of the labium, as described by Meinert (67, p. 369) and Grassi (’86°, 
p- 31), are probably homologous with the papille of Orchesella which I 
designated pip. (’99, Plate 3, Figure 24). 
For Lepisma, Heymons (97°, p. 590, Figur 11) gives, first, a pair of 
simple second maxillary fundaments and later (Taf. XXX., Figur 20) a 
long palpus with a small, basal, inner lobe, and states (p. 592) “Die 
Lobi oder spateren Ladentheile der Maxillen sind in diesen Stadien erst 
