FOLSOM: MOUTH-PARTS OF ANURIDA MARITIMA, 131 
als sehr kleine unscheinbare Vorspriinge erkennbar, welche medialwarts 
an der Basis der Taster hervorwuchern.” This is contrary to the condi- 
tions in Anurida, where the palpus is certainly itself an outgrowth from 
the siraple, primary papilla (Figure 13). Lepisma agrees with Anurida, 
however, in that the galea and lacinia are derived from the inner lobe of 
a biramous fundament (Heymons,’ 97°, Taf. XXX. Figur 17), and dis- 
agrees with Campodea if, in the latter genus, as Uzel implies, the galea 
buds from the palpus. The finished labium of Lepisma, as I shall show, 
is remarkably like that of Orthoptera. 
The labium of Machilis, as described by Oudemans (’88, pp. 185-186, 
Taf. II. Figuren 28, 29), resembles that of Campodea and Collembola in 
being deeply cleft, and having the salivary ducts opening in similar posi- 
tions, but it more nearly approaches the Orthopteran type in the position 
and structure of the terminal lobes, the mentum, and the three-jointed 
palpi. Each terminal lobe is subdivided into four lobes, which in all 
probability collectively represent galea and lacinia. 
Ayers (84, p. 241, Plate 18, Figures 20-22; Plate 19, Figure 5), as 
already quoted (p. 124), has traced the development of the second maxille 
of Gicanthus as far as the trilobed stage, stating the lobation to be more 
prominent in the second than in the first maxillary appendage. The 
fact that the second maxille of Anurida develop upon the Orthopteran 
type is important. In Lepisma, the trilobed fundaments agree with 
those of Orthoptera even as to the greater length of the palpus. 
In the finished labium of Cicanthus (Packard, ’83*, Plate XXVII. 
Figure 9) the derivatives of each trilobed fundament are easily identified 
as three-jointed palpus, galea, lacinia, palpifer, and mentum, — the last 
two structures having doubtless arisen from the common stalk, or stipes. 
Although the labium is constructed upon the same plan in all Orthoptera, 
we may best select Blatta for comparison with Lepisma. The agreement 
between Blatta (Mnhr, 777, Taf. II. Figur 11; Packard, ’83*, Plate 
XXVII. Figure 14) and Lepisma (Muhr, ’77, Taf. VIII. Figur 46; 
v. Stummer-Traunfels, ’91, Taf. II. Figur 17) is surprising. Galez and 
lacinize clearly correspond in the two, as do the mentum, palpifers, and 
palpi, the last, however, having three segments in Blatta and four in 
Lepisma. Muhr, in fact, included Lepisma among Orthoptera, as have 
some other authors. 
It is now agreed that the first and second maxille of Orthoptera are 
homodynamice, and, more inferentially, that the same is true of other in- 
sects. The exact agreement first recognized, according to Packard (98, 
p- 69), by Miall and Denny (’86), was detected long before, at least, by 
