142 PUCCINIA 



This species was formerly undistinguished from the numerous forms 

 on allied species of Composite, until Jacky proved experimentally (in 

 l.s!»:i tliai ii i- confined to the sub-genus Carduus, and could doI be 

 transferred to Cnicus or Cirsium. The uredo form described above (after 

 Sydow) is the secondary uredo ; the primary form, which does not always 

 occur, is described as ampbigenous and larger, often on the mid-ribs 

 2 — 4 mm. long, and remaining long covered by the epidermis. 



Distribution : Europe and Siberia. 



16. Puccinia Cardui-pycnocephali Syd. 



Puccinia Cardui-pycaocepltaU Sydow, Monogr. i. 34, f. 35. Massee, 

 Journ. Bot. xlvi. 152 ; Trans. Brit. Myc. Soc. (1909), iii. 123. 



Uredospores. Sori hypophyllous, without spots, scattered, 

 minute, pale-brown; spores globose or subglobose, very deli- 

 cately echinulate, pale-brown, 22 — 26 p,. 



Teleutospores. Sori similar, hidden in the tomentum of the 

 host, brown ; spores oblong, rounded above and more or less so 

 below, not thickened, hardly constricted, smooth, pale-brown, 

 38 — 50x16 — 23//,; epispore thin; pedicels thin, hyaline, up 

 to 40 \i long, but deciduous. 



On leaves and stems of Carduus pycnocephalus. Very rare; 

 Sidmouth (Dr Mayor), and between Eastbourne and Hastings 

 (G. Massee). Only recorded for Britain and Italy. July, 

 August. 



Distinguished from P. Carduorum by its longer teleutospores, which 

 are smooth, and not plainly verruculose as in that species. But sometimes 

 the spores are said to approach those of P. galatica Sydow, which occurs 

 on the same host in Asia Minor, in being smaller, darker-coloured, thick- 

 walled, and delicately punctate. According to Massee (I.e.) these are 

 rather two poles of one species than two distinct si>ecies. 



17. Puccinia Cirsii Lasch. 



Trichobasis Cirsii Lasch : Cooke, Micr. Fung. p. 224. 



Puccinia Cirsii Lasch, in Rab. Fung. Eur., no. 89. Cooke, Grevillea, 



iv. 109 ; Micr. Fung. p. 206. Sydow, Monogr. i. 55. Fischer, 



Ured. Schweiz, p. 217, f. 168 — 171. 



Uredospores. Sori epiphyllous or hypophyllous, on paler 

 spots, scattered, minute, girt by the epidermis, pulverulent, cin- 

 namon ; spores globose to ovate, echinulate, pale chestnut-brown, 



