ON CAMPANULACE^E 159 



cinnamon ; spores globose to ellipsoid, echinulate, yellow-brown, 

 24 — 29 x 16 — 25 /j,, with two germ-pores. 



Teleutospores. ISori similar and often on the stems, but 

 dark-brown; spores ellipsoid or somewhat 

 ovate, rounded and not thickened above, 

 scarcely constricted, usually rounded 

 below, very delicately verruculose, brown, 

 25 — 40 x 16 — 24 //, ; epispore thin ; pedi- 

 eels hyaline, usually very short. Fig uo p ffimu; .. 



On leaves and stalks of Hieracium, Teleutospores. 



H. boreale, H. murorum, H. Pilosella, II. umbellatum and 

 various subspecies. Very common. May — November. (Fig. 

 110.) 



It was proved by Jacky that this species, which occurs so abundantly 

 on Hieracium, cannot be transferred to other genera of Compositse. As a 

 similar fact has been demonstrated for many other species of Uredinales, 

 there is sufficient ground for the assumption, now generally made, that 

 most species of Puccinia, etc., which are parasitic on different genera 

 should be regarded as distinct, even when no experimental evidence exists 

 in favour of that course. Jacky was also inclined to suspect that 

 P. Hieracii might hereafter be divisible into a number of biologic races, of 

 which, however, he only indicated one, that on H. villosum belonging to 

 the section Pilosella. Rene Probst (Centralbl. f. Bakt. 1909, 2. xxii. 676) 

 not only divided P. Hieracii into 13 biologic races, arranged under two 

 subspecies, P. Piloselloidarum on the section containing H. Pilosella and 

 its allies, and P. Hieracii (sens, strict.) on the other species, forming the 

 section Euhieracium, — but he goes on to reduce the question of such 

 races to an absurdity by "proving" that one of them was restricted to a 

 mere form of a variety of a subspecies {Hieracium Pilosella, subsp. 

 vulgare, var. gemdnum, forma subpilosum). 



The two subspecies may perhaps be defensible, if they are distinguished 

 morphologically, as Probst states, by the fact that in P. Piloselloidarum 

 the germ-pores of the uredospore are strictly equatorial, but in P. 

 Hieracii they are removed towards the upper pole. 



Distribution : Europe, Asia Minor, North America, Chili. 



33. Puccinia Campanula? Carm. 



Puccinia Campanulae Carmich. in Berk. Engl. Fl. v. 365. Cooke, 

 Handb. p. 498 ; Micr. Fung. p. 205. Plowr. Ured. p. 200. Sacc. 

 Syll. vii. 677. Sydow, Monogr. i. 196, f. 182. Fischer, Ured. 

 Schweiz, p. 175, f. 136. 



