ON POLYGONACE/E 225 



95. Puccinia Conopodii-Bistortae Kleb. 



Puccinia Bistortae DC. Flor. fr. vi. 61. Cooke, Micr. Fung. p. 204 ; 



Grevillea, ii. 161. Plowr. Ured. p. 192 (?). Sydow, Monogr. 



i. 571 (all pro parte). Soppitt, Grevillea, xxii. 45 ; Card. Chron. 



1895, xviii. 773. 

 P. Conopodii-Bistortae Kleb. Wirtsw. Rostp. p. 318. 



^Kcidivspores. /Ecidia on large swellings of a bright orange 

 colour, immersed, spherical or flat, not at all cup-shaped, margin 

 not projecting; spores delicately verruculose, orange, 15 — 20 pu. 



Uredospores. Sori hypophyllous, minute, roundish, yellowish- 

 red, soon naked; spores globose to shortly ellipsoid, finely 

 echinulate, pale yellowish-brown, 21 — 24 p,. 



Telmtospores. Sori hypophyllous, scattered or united in 

 roundish groups, soon naked and pulverulent, dark- 

 brown ; spores oblong or subclavate, rounded at 

 both ends or slightly oblique at the apex, faintly 

 constricted, clear yellowish-brown, 28 — 42 x 16 — 

 25 p, ; epispore equally thick, smooth, but some- 

 times marked with a very few longitudinal or m ^ 

 oblique rows of delicate warts ; no papilla on P. Conopodii- 

 the germ-pores ; pedicels hyaline, short, very Teleutospoi-e. 

 deciduous. 



iEcidia on Conopodium denudatum, Yorkshire, May; teleuto- 

 spores on Polygonum Bistorta, June — August, not common. 

 (Fig. 173.) 



The Puccinias on Pol. Bistorta are not yet well known. There appear to 

 be at least four (or five) distinct forms, which are divisible morphologically 

 into two groups — (1) those whose teleutospores have no apical papilla, 

 (2) those which have a small hemispherical papilla at the summit ; the 

 latter are called by Sydow in the Monographia P. mammillata Schrot. 

 (Pilz. Schles. p. 340). They are all heteroscious. 



Of the former group, one form has an ajcidium on Conopodium, and its 

 other spore-stages on P. Bistorta. It is the one mentioned above ; the life- 

 history was first demonstrated by Soppitt (I. c). It is remarkable as 

 being the first instance known of a hetercecious Puccinia that had its 

 teleutospores on a Dicotyledon. The second form has its secidium on 

 Angelica silvestris and Carum Carui and its teleutospores on Pol. Bistorta 

 and Pol. cieiparum. It would perhaps be better divided into P. Angelicae- 

 Bistortae Kleb. (= P. Car i- Bistortae) and P. Polygoni-vivipari Karst. The 

 connection in the former has been demonstrated by Klebahn and Fischer ; 



o. u. 15 



