OX GRAMINE.K 255 



species and is accompanied by few uredospores, while that on Lolium 

 perenne (P. Lolii) occurs only in the autumn with a profuse simultaneous 

 development of uredospores. This agrees with my experience. Never- 

 theless, I consider that P. coronata and P. Lolii are merely biological 

 forms of one species. The experiments of Carleton, in the United State:-, 

 have shown that the distinctions of hosts are quite insufficient to dis- 

 criminate the two forms ; among others he succeeded in infecting a host, 

 stated by Eriksson to be confined to P. coronata, with spores from a host 

 belonging to P. Lolii. 



P. Festucae Plowr. belongs to the same group, but has its secidium on 

 Honeysuckle. Crown Rusts have been found in Europe on many other 

 species and genera of grasses (Sydow, i. 705), but they cannot, in the 

 absence of cultures, be even temporarily arranged under the two heads. 

 Barclay described a form from Simla (P. coronata var. himalensis) on 

 Brachy podium silvaticum with its secidium on Rhamnus dahurica ; this 

 has since been raised by Dietel to a species (P. himalensis). 



It is a remarkable fact that the only other species of Puccinia known, 

 provided with the same processes, are two (P. Mesnieriana = digitata, and 

 P. ScJiweinfurthii) whose telcutospores occur on Rhamnus. A somewhat 

 similar form, but with much longer apical processes, is found on Lonicera 

 in Turkestan (P. longirostris Kom.). These instances are similar to those 

 of P. fusca and P. Pruni-spinosae already mentioned and can be explained 

 in the same way ; see Grove, New Phytologist, 1913, p. 89. 



119. Puccinia Lolii Nielsen. 



JEcidium crassum Pers. ; Cooke, Handb. p. 538 ; Micr. Fung. p. 196 p.p. 



jE. Cathartici Schum. PL Sail. ii. 2:25. 



Puccinia coronata Corda ; Cooke, Handb. p. 494 ; Micr. Fung. 



p. 203 p.p. Plowr. Ured. p. 163 p.p. Sacc. Syll. \ ii. 623 p.p. 

 P. Lolii Niels. Ug. for Landrnaend. i. 549 (1875). Sydow, Monogr. i. 



704. McAlpine, Rusts of Australia, p. 123 and many figures. 

 /'. coronifera Kleb. Zeitsch. f. Pflanzenkr. iv. 132 (1894). Fischer, 



Ured. Schweiz, p. 375, f. 271. 



jEcidiospores. yEcidia hypophyllous or on the petioles, 

 seated on yellow or purplish spots, scattered or arranged in 

 groups, producing distortion of the parts, cylindrical, with a 

 whitish torn revolute margin ; spores very delicately verrucu- 

 lose, orange, 1G — 25 x 12 — 20 ft. 



Uredospores. Sori amphigenous, scattered or in large 

 patches, forming blister-like swellings, minute, sometimes con- 

 fluent, lanceolate or more or less oblong, pulverulent, orange ; 

 spores globose to obovate, echinulate, yellow, 18 — 27 x 16 — 24/a, 



