12 ILLINOIS BIOLOGICAL MONOGRAPHS [480 



erroneous and that of Ellis and Martin as correct and to recognize M. 

 puiggarii on Rubus, as an entirely different species. 



In the head cell of each capitate hyphopodium there is a small, clear 

 spot, about 3/A in diameter, which appears to be a hole in the lower 

 side of the hyphopodium, and if so probably indicates the breaking off 

 of a haustorium. Similar marks are found upon the capitate hyphopo- 

 dia in many species but they are not usually so conspicuous as in this 

 species. 



3. M. GUiGNARDi Gain. 



Bull. Soc. Myc. de France 8 : 176, 1892. 

 On Turpinia panniculata Vent. Fig. 3. 



Maricao, July 19, '15, 8922, Sept. 20, '13, 3685. 



This differs somewhat from the type as described by Gaillard in its 

 smaller perithecia, about 300//., shorter perithecial setae, about 100- 

 150/i, and smaller ascospores, 54 x IS^u,. 



It is of special interest since it is the only species known which 

 has a 3-septate spore, without mycelial setae, and with perithecial setae. 

 The hyphopodia are very characteristic and agree well with the figures 

 of Gaillard. The type of the species was from Ecuador on host unknown. 



SECTION B 



Spores 4-septate, no setae or appendages 



This section corresponds to the Section I C. a. of Gaillard without 

 the forms with perithecial appendages. As represented in Porto Rico 

 it presents many puzzling aspects, occurring on a large number of 

 hosts of many families and presenting variations which though easy 

 to recognize are difficult to define with accuracy. The spores in all are 

 quite uniform and neither in size, shape nor constriction lend themselves 

 to description. There are no perithecial or mycelial setae. Therefore 

 the perithecium, the mycelium and hyphopodia must furnish the dis- 

 tinctions. Moreover, such variations as do occur in the perithecium are 

 minor and difficult to state definitely. For this reason many students 

 have referred such forms on numerous hosts to M. glabra. A study of 

 the Porto Rican material both in the field and under the microscope 

 convinces me that many of these forms which may agree in spores and 

 perithecia and which show variation in general habit in mycelium and 

 hyphopodia are really distinct species. Plants heavily infected with 

 one of these forms of Meliola may be found growing among plants sus- 

 ceptible to another form, but which are entirely devoid of Meliola. 



In the treatment of the Porto Rican species I have removed from 

 M. glabra all which it seems very evident are not co-specific with the 



