210 WHEELER. 



of Eusphinctus steinhcili Forel (Fig. 2). It is very long and slender, 

 non-tubereulate and covered with bifurcate hairs. No pupte of Cera- 

 pachyini have been seen, so that we are unable to say whether they 

 are naked or enclosed in cocoons like the pupae of all other Ponerinse, 

 except those of the genus Disrofhi/rca. 



Emery in the "Genera Insectorum" recognizes four genera of 

 Cerapachyini, all of which are represented in Australia, namel\', 

 Spkinctomyrmcx , Cerapackys, Phyracaccs and Lioponcra, the first 

 comprising the subgenera Sphinctoun/rmc.v s. str. and Eusphinciws, 

 the second four subgenera: Ccrapaclu/s s. str., Parasyscia, Ooccra'a 

 and Syscia. Of the latter only Syscia is known to be represented in 

 Australia. The taxonomic status of some of these groups is still 

 doubtful, owing largely to incomplete information concerning their 

 sexual phases. Emery (Zool. Jahrb. Abt. Syst. 8, 1895, p. 693, PI. 

 14, Fig. 4) described and figured the genitalia of a male of an unde- 

 termined species of Eusphinctus, presumably from the Indomalayan 

 region. He found that the inner paramera terminate as vertical 

 plates with dentate inferior borders, and that the vosellse are movably 

 articulated, well differentiated and bear a vestigial lacinia at their 

 bases. Of the males of the Australian species which have been in- 

 cluded in the genus Sphinctomyrmc.v, nothing is known. Santschi 

 has recently described a male ant from Africa as S. rufiventris, but 

 its generic status seems to me to be open to doubt. The peculiarities 

 of the females of the Australian forms have, in my opinion, an impor- 

 tant taxonomic bearing, as will be evident from the following considera- 

 tions. 



The genus Sphinctoviyrmcx was erected by Mayr (Verb. zool. bot. 

 Ges. Wien 16, 1866, p. 895, PL 20, Fig. 8) for the accommodation of 

 a single dealated female specimen described from Brazil as S. stdli. 

 During half a century no one has succeeded in again finding this 

 insect. Mayr's figure shows that its thorax has well de\eIoped 

 sclerites essentially like those in ordinary female ants. The antenntie 

 are 12-jointed, the pygidium emarginate. In 1895 Emery erected 

 a genus Eusphinctus for a worker ant (E. furcatus) from Lower Bur- 

 mah, with 11-jointed antennje, but with the pygidium emarginate 

 and the gastric segments separated from one another by deep con- 

 strictions as in the Brazilian stdli. In 1897 a second species {E. 

 cribratus) was described by Emery from New Guinea and Forel added 

 a third from Bengal {E. taylori) in 1900. Only workers of these forms 

 have come to light. In the meantime Forel received several species 

 from Australia, some of which had 11-jointed, while others had 12- 



