FLEMINGER: INTEGUMENTAL ORGANS IN GENUS EUCALANUS 



Types (uid type locality: Holotype: 9.USNM 

 143844. Cruise Downwind, stn 23a. 45-cm net, 

 0-352 m; 19 Oct. 1957, 0000-0100 h; 38°46'S, 

 83°20'W. 



Reference: 3 99. USNM 143845. Cmise Down- 

 wind, stn 28, 1-m net, 0-283 m; 31 Dec. 1957, 

 0520-0542 h; 27°08.5'S, 72°02'W. 



3 99. USNM 143846. Cruise Atlantis 11-13, 

 stn 8, %-m net, depth? 5 Oct. 1964, 1410-1435 

 h; 41°26'N,55°45'W. 



1 9. USNM 143847. Cruise Dodo VI, stn 55, 

 1-m net, 0-200 MWO; 22 Aug. 1964, 1845-1858 

 h; 10°14'N,51°19'E. 



Remarks: Brodsky's (1962, p. 113) record of 

 (?) attenuatus appears from his illustration 

 (Figure 14) and the locality off northern Japan 

 to be referable to sewelli. Other records that 

 appear to be assignable to sewelli include Tanaka 

 (1956, p. 266, Figure 4a), Ramirez (1969), and 

 Roe (1972). This species is named to honor R. B. 

 Seymour Sewell who among his many signifi- 

 cant contributions to the copepod literature 

 inspired study of their integumental organs 

 and directed attention to the complexity exist- 

 ing in the attenuatus group. 



Encalauiis parki, n. sp. 



(Figures Is, 15a, 18e, f, 1, m s, t) 



E. attenuatus: Park 1968, p. 533, PI. 1, Figs. 

 16, 17, PI. 2, Figs. 1-16, PI. 3, Figs. 1-13, 

 large form only. 



Diagiiosis: Adult female: Mandibular basis 

 with four setae. Sites of integumental organs 

 similar to those of seivelli except for presence 

 of one medial tergal site on both the genital 

 segment and on Abd.III, one or more pleural 

 sites on Abd.IV-V, and only one transverse 

 tergal row on Abd.IV-V (Figure 15a). 



Adult male: Forehead prominent, protruding 

 as an elongate triangle about twice as far be- 

 yond base of rostrum as in seivelli and attenu- 

 atus (Figure 18f). Integumental organs on 

 cephalosome and thorax as in female; Abd.II 

 with or without one tergal site (in 11 specimens, 

 6 with and 5 without). Mandibular basis usually 

 with four setae, one specimen in 11 with three 

 setae. 



Additional description: Adult female with 

 conical forehead and rounded apex as in Figure 

 18e, differs from typical acutely pointed fore- 

 head of sewelli. 



Total length: Adult female: range 6.40 to 7.17 

 mm, mean = 6.744 mm, s = 0.1982, N = 18 

 (Figure 16). 



Adult male: range 5.20 to 5.96 mm, mean = 

 5.620 mm, s = 0.2380, A' = 10. Localities of 

 specimens listed in Table 1. pooled sample 

 represents all adults on hand. 



Geographical distribution: All records (Fig- 

 ure 17) are from the North Pacific Ocean. Most 

 fall within the Transition zone (Brinton, 1962, 

 p. 202) but a number also appear along long. 

 155°W extending south to lat. 32°N, following 

 the line of division between western and eastern 

 North Pacific Central Water (Sverdrup, Flem- 

 ing, and Johnson, 1942, Figure 209A) which is 

 supported by the relative elevation of the iso- 

 therms in the upper 200 m of the mid-latitudes 

 in the North Pacific relative to those of the 

 South Pacific (Reid, 1965, Figure 2). 



Types and type locality: Holotype: 9. USNM 



143841. Cruise Norpac H5508. stn 24-42, 1-m 

 net, 0-200 MWO; 21 Aug. 1955; 1556-1610 

 PST; 40°26'N, 146°37'W. 



Reference: 1 9, specimen no. 2. USNM 



143842. Cruise Norpac H5508, stn 24 (up), 1-m 

 net, 0-200 MWO; 25 Aug. 1955, 1620 PST; 

 42°17'N, 146°16'W. 



1 9, specimen no. 1. USNM 143843. Cruise 

 Norpac H5508, stn 25 (up), 1-m net, 0-200 

 MWO; 25 Aug. 1955, 1816-1831 PST; 42°17'N, 

 150°00'W. 



Remarks: Large forms of attenuatus s.l. have 

 been noted from the North Pacific by several 

 authors (e.g., Tanaka, 1956, p. 266, Figure 4b; 

 Lang, 1965 Park, 1968). Park (1968) fully 

 appreciating its distinctiveness described it in 

 detail. He relinquished his original intention of 

 naming it as a distinct species upon my recom- 

 mendation, which was based on the apparently 

 extensive variability generally thought to pre- 

 vail in Eucalanus. At that time there was no 

 reliable basis for distinguishing genetic from 

 phenetic variation and reliable capture records 



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