U.S. DEPARTMENT OF COMMERCE 



Maurice H. Stans, Secretary 

 NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION 



NATIONAL MARINE FISHERIES SERVICE 

 Philip M. Roedel, Director 



FISHERY BULLETIN 



The Fishery Bulletin carries technical reports on investigations in fishery science. The Bulletin of the 

 United States Fish Commission was begun in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 

 and the Fishery Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents through 

 volume 46; the last document was No. 1103. Beginning with volume 47 in 1931 and continuing through volume 

 62 in 1963, each separate appeared as a numbered bulletin. A new system began in 1963 with volume 63 in which 

 papers are bound together in a single issue of the bulletin instead of being issued individually. 



Bulletins are distributed free to libraries, research institutions, State agencies, and scientists. Some Bulle- 

 tins are for sale by the Superintendent of Documents, U.S. Government Printing Office, Washington, D.C. 20402. 



EDITOR 



Dr. Reuben Lasker 



Scientific Editor, Fishery Bulletin 



National Marine Fisheries Service 



Fishery-Oceanography Center 

 La Jolla, California 92037 



Editorial Committee 



Dr. Elbert H. Ahlstrom 



National Marine Fisheries Service 



Dr. William H. Bayliff 



Inter-American Tropical Tuna Commission 



Dr. Daniel M. Cohen 



National Marine Fisheries Service 



Dr. Howard M. P'eder 

 University of Alaska 



Mr. .John E. Fitch 



California Department of Fish and Game 



Dr. Marvin D. Grosslein 

 National Marine Fisheries Service 



Dr. J. Frank Hebard 



National Marine Fisheries Service 



Dr. John R. Hunter 



National Marine Fisheries Service 



Mr. John C. Marr 



Food and Agriculture Organization of 

 the United Nations 



Dr. Arthur S. Merrill 



National Marine Fisheries Service 



Dr. Virgil J. Norton 

 University of Rhode Island 



Mr. Alonzo T. Pruter 



National Marine Fisheries Service 



Dr. Theodore R. Rice 



National Marine Fisheries Service 



Dr. Brian J. Rothschild 

 University of Washington 



Dr. Oscar E. Sette 



National Marine Fisheries Service 



Mr. Maurice E. Stansby 

 National Marine Fisheries Service 



Dr. Majmard A. Steinberg 

 National Marine Fisheries Service 



Dr. Roland L. Wigley 



National Marine Fisheries Service 



