Figure 48. — Palaemonetes {Palaemonetes) intermedins 

 Holthuis. A, anterior part of body in lateral view; B, 

 antennule ; C, antennal scale ; D, second leg of female ; 

 E, fingers of second leg of female; F, third leg; A-F 

 X 4.75 (after Holthuis, 1952). 



dorsal margin up to often bifurcate tip, distal 

 teeth more widely spaced than proximals; lower 

 margin with 4 or 5 (occasionally 3) teeth. Cara- 

 pace smooth; antennal spine present; branchioste- 

 gal spine on anterior margin just below bran- 

 chiostegal groove. Eyes well developed. Anten- 

 nular peduncle as in vulgaris, basal article with 

 slender stylocerite reaching about to middle of 

 article, anterolateral spine of article strong, over- 

 reaching rounded anterior margin; upper anten- 

 nular flagellum with both rami fused for 7 to 

 10 joints; free part of shorter ramus with 7 to 

 12 joints, longer than fused portion. Antennal 

 scale slender, 3 to nearly 4 times as long as broad 

 in females, even more slender in males; outer 

 margin straight or slightly concave; terminal 

 tooth reaching about to end of lamella. 



First legs almost reaching tip of antennal scale ; 

 fingers as long as palm; carpus twice length of 

 chela and slightly longer than merus. Second legs 

 in adult female usually with almost entire chela 

 reaching beyond antennal scale; fingers a little 

 over half length of palm, cutting edge of dactyl 

 with one proximal tooth, remainder of cutting 

 edges of both fingers entire; carpus 1.2 to 1.5 

 times length of palm and as long as merus. Sec- 

 ond leg of male somewhat more slender than in 

 female ; only fingers reaching beyond scale ; carpus 



as long as merus. Third leg with propodus less 

 than twice as long as carpus. Fifth leg with 

 propodus about three times as long as dactyl, 

 twice as long as carpus. 



Abdomen smooth; pleura of fifth segment with 

 tip rectangular or slightly acute; sixth segment 

 1.5 times length of fifth, somewhat shorter than 

 telson. Telson with two pairs of dorsal spines; 

 anterior pair somewhat behind middle; second 

 pair halfway between these and tip; posterior 

 margin with strong median point flanked by two 

 pairs of spines, inner pair longest and between 

 these, two feathered setae. Outer margin of 

 uropodal exopod with a strong terminal tooth 

 flanked by a slender movable spine medially. 



Measurements. — Length of body : male, 30 mm. ; 

 ovigerous females, 20 to 42 mm. 



Variations. — In males and juveniles, the legs 

 reach less far forward than in ovigerous females 

 (Holthuis, 1952). The second chelae of some fe- 

 males have one tooth on the cutting edge of each 

 finger. 



Color. — Transparent in life. 



Habitat. — Estuarine waters, especially in beds 

 of submerged vegetation. 



Type locality. — Iron Box Bay, Chincoteague 

 Bay, Va. 



Known range. — Long Island, N.Y., to Port 

 Aransas, Tex. (from specimens examined by 

 Holthuis, 1952). Literature records: Vineyard 

 Sound to Aransas National Wildlife Refuge, Tex. 

 (Holthuis, 1952). 



Remarks. — The confused taxonomic status of 

 this species in literature is discussed in the account 

 for P. vulgaris and dealt with in more detail by 

 Holthuis (1952). Ovigerous females have been 

 found from February to April in Texas (Hedg- 

 peth, 1950), and from May to September in 

 Virginia and the Carolinas. 



Palaemonetes (Palaemonetes) pugio Holthuis 



Figure 49 



Palaemonetes vulgaris: Hay and Shore, 1918, p. 393, pi. 27, 

 fig. 5. 



Palaemonetes {Palaemonetes) pugio Holthuis, 1949, p. 95, figs. 

 2, m-o.^Holthuls, 1952, p. 244, pi. 55, figs, g-1 (rev.). 



Recognition characters.- — Rostrum reaching to 

 or slightly beyond end of antennal scale ; straight, 

 sometimes slightly upturned at tip; dorsal mar- 

 gin with 7 to 10 (usually 8 or 9) teeth, distal teeth 

 more widely spaced than proximal teeth, first 



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