center of upper surface of each cheliped; re- 

 mainder of body light gray-tan; eyestalks brown 

 below cornea; hairs on antennules rusty brown 

 (Provenzano, 1959, for west Florida form). 



Habitat. — Deep channels of harbors and littoral 

 waters; also found in shallow estuaries near 

 ocean; near low-tide mark to 25 fathoms. 



Type locality. — [East] coast of United States. 



Known range. — Vineyard Sound, Mass., to 

 northeastern Florida; Key West, Fla., to Texas 

 (Provenzano, 1959). 



Remarks. — Blake (1953) reported P. poUicaris 

 from the Pleistocene of Maryland. 



Ovigerous females have been reported in March 

 from Florida (Provenzano, 1959). They have 

 been found in January and November in North 

 Carolina, and in April in Chesapeake Bay. 



Pagurus impressus (Benedict) 



Figure 104 



Eupagurus impressus Bendtct,,1892, p. 5. 



Pagurus impressus: Provenzano, 1959, p. 399, fig. 15 (rev.). 



Recognition characters. — Anterior shield of 

 carapace about as broad as long, flattened. Ros- 

 trum much rounded, in line with somewhat more 

 angular lateral projections of front. Eyestalks 

 slender, about three times longer than greatest 

 width, cornea dilated and flattened ; eye scales with 

 moderately slender, acuminate, slightly excavated 

 anterior lobe, subterminal spine large. Antennular 

 peduncles exceeding eyestalks by at least half of 

 last article. Antennal peduncles slightly exceed- 

 ing cornea ; acicles curving outward, reaching to 

 base of cornea, hairy on medial edge. 



Chelipeds unequal, right much larger than left, 

 upper surfaces dented, both covered with small, 

 closely crowded granules, outer margins bearing 

 enlarged granules or small spines. Dactyl of ma- 

 jor cheliped with a rounded angle on medial bor- 

 der near tip followed by marginal spines or gran- 

 ules, tips of fingers acuminate; carpus with five 

 longitudinal rows of small spines often obscured 

 by irregularly arranged additional spines, a row 

 of well-developed spines along inner edge. Minor 

 cheliped reaching to angle of major dactyl; a row 

 of spines along inner margin of hand and carpus; 

 fingers somewhat spooned at tips, cutting edges 

 well defined, tips corneous, dactyl with medial 

 border tuberculate; carpus with several rows of 

 small spines. First and second walking legs with 



Figure 104. — Pagurus impressus (Benedict). Anterior 

 part of body in dorsal view, approximately X 2.5 (after 

 Provenzano, 1959). 



dactyls much longer than propodi ; first pair with 

 row of small, well-defined spines along upper mar- 

 gin of propodus and carpus; second pair with 

 spines reduced. Third legs reduced, fourth re- 

 duced and turned on back. 



Measurements. — Length of carapace: male, 25 

 mm.; female, 16 mm. (Provenzano, 1959). 



Color. — Hands solid rust to chocolate brown, 

 other appendages brownish with alternating thin 

 bands of light color; antennae, antennules, and 

 peduncles yellow; eyestalks brown above, bright 

 blue below with bright scarlet at base of eyestalks 

 (Provenzano, 1959). 



MARINE DECAPOD CRUSTACEANS OF THE CAROLINAS 



129 



