Figure 139. — Hepatus pudibundus (Herbst) . A, male first 

 pleopod ; B, male second pleopod ; A-B approximately 

 X 8 ; C, abdomen of male ; D, abdomen of female ; C-D 

 approximately X 2 (after Holthuis, 1959). 



Habitat. — Beach to 6 fathoms. 



Type locality. — Martinique. 



Known range. — Georgia to State of Sao Paulo, 

 Brazil. 



Remarks. — Holthuis (1959) reported an ovig- 

 erous female in April from Surinam. Several of 

 the specimens Holthuis studied carried one or 

 more sea anemones on the carapace and one bore 

 barnacles. 



Hepatus epheliticus (Linnaeus) Calico crab, Dolly Varden 

 crab 



Figure 140 



Cancer epheliticus Linnaeus. 1763, p .414. 



Hepatus epheliticus: Hay and Shore, 1918, p. 422, pi. 37, 

 flg. 1. — Rathbun, 1937, p. 238, pi. 70, figs. 3-4; pi. 71, figs. 1-4 

 (rev.). 



Recognition characters. — Carapace covered 

 with large spots, about two-thirds as long as wide, 

 convex above, regularly arcuate in front, strongly 

 narrowed behind, almost smooth, with indistinct 

 lines of low granules on gastric region and pos- 

 terior part of branchial region. Front narrow, 

 truncate, tuberculate, not dentate, and placed 

 higher than continuation of anterolateral borders 

 of carapace; anterolateral borders dentate with 

 denticles more prominent than in H. pudibundus. 

 middle denticle of each tooth projecting. 



Chelipeds moderately strong; carpus and hand 

 with lines of coarse tubercles on outer face and a 



three- to four-toothed crest on superior margin of 

 hand. Dactyls of walking legs with a coating of 

 short, dense pubescence except for tip and a nar- 

 row smooth line on each side. 



Measurements. — Carapace of male: length, 46 

 mm. ; width, 67 mm. 



Variations. — In a series representing various 

 ages a great deal of variation is shown, the granu- 

 lations being relatively much coarser and the 

 spots more numerous and variable in color in 

 juveniles. 



Color. — Dark gray or brownish with numerous, 

 rather large, round or irregular spots of light red 

 (muddy lavender to light red in young) with 

 darker borders scattered over carapace. 



Habitat. — This species is often found in depths 

 of a few fathoms in channels of Beaufort, N.C., 

 harbor, though such individuals are not so numer- 

 ous or large as those taken in the ocean outside. 

 Gunter (1950) found the species in water ranging 

 from 28.7 to 35.2 % . Two to 25 fathoms. 



Type locality. — Carolina. 



Known range. — Chesapeake Bay to Gulf of 

 Campeche, Mexico; Cuba; Jamaica; Dominican 

 Republic. 



Remarks. — Though recorded in depths to 25 

 fathoms, Hildebrand (1954, 1955) found this spe- 

 cies most common shoreward of 12 fathoms in 



Figure 140. — Hepatus epheliticus (Linnaeus). Male in 

 dorsal view, legs of left side not shown, 20 mm. indi- 

 cated. 



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