2. Hepatic carina prominent 3 List of Species 



Hepatic carina absent or ill defined p (p ) 7?!<3?iWo/i. Fabricius and P. {P.) escu- 



Fenneropenaeus Jcntus Haswell in the Indo-Pacific; and P. (P.) 



3. Thelycum open type. Petasma with ventral se»iis(//ea-iMs de Haan in the Indo-Pacific and east- 



costa short, not reaching distal margin of ern Mediterranean Sea, 



lateral lobe Litopenaeus 



Thelycum closed type. Petasma with ventral SUBGENUS Fenneropenaeus New Subgenus 



costa long, reaching distal margin of lateral rj,^^ species, Penaem {F e7ineropenaeus) in- 



lobe Penmus s.s. ^^^.^^^^ jj j^-j^^^ Edwards, here designated. 



Subgenus Litopenaeus New Subgenus Diagnosis 



Type species, Penaeu.s {Litopenaeus) vawmrmi thelycum of closed type. Petasma with ventral 



Boone, here designated. ^^^^^^ ^^^^^ strongly curved distally, and with 



Diagnosis median lobes only slightly projected distally. Ad- 

 rostral sulcus and carina short, extending about to 



Thelycum of open type, lacking sperm recepta- ig^.gj ^f epigastric tooth, sulcus shallow and carina 



cle. Petasma formed by relatively simple petasmal attenuated posteriorly; gastrofrontal carina ab- 



endopods, with short ventral costae; median lobes ggj^^. hepatic carina absent or ill defined; gastro- 



of petasma not extended distally, not or just barely orbital carina short or absent, if present extendmg 



reaching distal margin of lateral lobes. Carapace anteriorly not more than two-thirds, or occupying 



with adrostral carina and sulcus short, extending middle one-third of the distance from hepatic 



to, or only slightly posterior to, epigastric gpine to postorbital margin, 

 tooth; gastrofrontal carina absent; hepatic carina 



well developed; gastro-orbital carina short, ex- Etymology 



tending anteriorly not more than three-quarters, iT'enwerojsewa^MS for Fenner A. Chace, Jr., whose 



usually two-thirds, of the distance from hepatic advice through the years has been very helpful to 



spine to postorbit al margin. ^^^^  ^^ ^^^ studies. 



Etymology List of Species 



Litoperuiem, from the Greek litos, meaning P. (7^.) wi^zVm.s H. Milne Edwards, P(/^.) me7'- 



pjj^jj^_ guiensh cle Man, P. {F .) penicillatus Alcock, and 



,. ,„ . P. (Z^.) omriteZwKisliinouye. The four species are 



L/lst ot opecies .^ 



Indo-Pacitic. 



P. [L.) vannamei Boone, P. (Z.) stylirostris SUBGENUS Melicertus 

 Stimpson, and P. (L.) occidentalis Streets in the 



eastern Pacific; P. (L.) setiferus (L.), and Type species, by monotypy, yl/eZ/ce/'^ws ^-/p-Wnw.? 



P. (Z.) schmitti Burkenroad in the western Rafinesque, 1814: 22 ( = 6'ancer A;era^/(«7'M.s For- 



Atlantic. skal, 1775). 



Subgenus Penaeu* S.S. Salamhria "Burkenroad"' Balss, 1957: 1518 



(nomen nudum) . 



Type species, Peruteu.^ (Penaeus) monodon Diagnosis 



Fabricius. ^ .,, , , 



. Thelycum of closed type. Petasma with ventral 



Diagnosis co%ti\& long, strongly curved or almost straight 



Thelycum of closed type. Petasma with ventral distally, and with distomedian projections usu- 



costae long, strongly curved distally, and with ally prominent. Carapace with adrostral carina 



median lobes only slightly prolonged distally. and sulcus long, extending posteriorly consider- 



Carapace with adrostral carina and adrostral sul- ably beyond epigastric tooth, usually almost reach- 



cus sliort, extending about to level of epigastric ing posterior margin of carapace; gastrofrontal J 



tooth; gastrofrontal carina absent; hepatic carina carina present; hepatic carina prominent; gas- 1 



well defined and gastro-orbital carina short, ex- tro-orbital carina long, extending anteriorly at 



tending anteriorly not more than two-thirds of the least, usually more than three-quarters of the dis- 



distance from hepatic spine to postorbital margin. tance from hepatic spine to postorbital margin. 



466 U.S. FISH AND WILDLIFE SERVICE 



