U.S. DEPARTMENTOFCOMMERCE 



Frederick B. Dent, Secretary 



NATIONALOCEANIC AND ATMOSPHERIC ADMINISTRATION 



Robert M. White, Administrator 



NATIONALMARINE FISHERIES SERVICE 

 Robert W. Schoning, Director 



Fishery Bulletin 



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The Fishery Biillciln carries Driginal research reports and technical notes on investigations in fishery science, 

 engineering, and economics. The Bulletin of the United States Fish Commission was begun in 1881; it became the 

 Bulletin of the Bureau of Fisheries in 1904 and the Fishery Bulletin of the Fish and Wildlife Service in 1941. Separates 

 were issued as documents through volume 46; the last document was No. 1103. Beginning with volume 47 in 1931 and 

 continuing through volume 62 in 1963, each separate appeared as a numbered bulletin. A new system began in 1963 

 with volume 63 in which papers are bound together in a single issue of the bulletin instead of being issued individually. 

 Beginning with volume 70, number 1. January 1972. the Fishery Bullciin became a periodical, issued quarterly. In this 

 form, it is available by subscription from the Superintendent of Documents, U.S. Government Printing Office, Washington, 

 D.C. 20402. It is also available free in limited numbers to libraries, research institutions. State and Federal agencies, 

 and in exchange for other scientific publications. 



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EDITOR 



Dr. Reuben Lasker 



Scientific Editor, Fishery Bulletin 



National Marine Fisheries Service 



Southwest Fisheries Center 

 La Jolla, California 92037 



Editorial Committee 



Dr. Elbert H. Ahlstrom 



National Marine Fisheries Service 



Dr. William H. Bayliff 



Inter-American Tropical Tuna Commission 



Dr. Daniel M. Cohen 



National Marine Fisheries Service 



Dr. Howard M. Feder 

 University of Alaska 



Mr. John E. Fitch 



California Department of Fish and Game 



Dr. Marvin D. Grosslein 

 National Marine Fisheries Service 



Dr. J. Frank Hebard 



National Marine Fisheries Service 



Dr. John R. Hunter 



National Marine Fisheries Service 



Dr. Arthur S. Merrill 



National Marine Fisheries Service 



Dr. Virgil J. Norton 

 University of Rhode Island 



Mr. Alonzo T. Pruter 



National Marine Fisheries Service 



Dr. Theodore R. Rice 



National Marine Fisheries Service 



Dr. Brian J. Rothschild 



National Marine Fisheries Service 



Mr. Maurice E. Stansby 

 National Marine Fisheries Service 



Dr. Maynard A. Steinberg 

 National Marine Fisheries Service 



Dr. Roland L. Wigley 



National Marine Fisheries Service 



Kiyoshi G. Fukano, Managing- Editor 



The Secretary of Commerce has determined that the publication of this periodical is necessary in the transaction of 

 the public business required by law of this Department. Use of funds for printing of this periodical has been approved 

 by the Director of the Office of Management and Budget through May 31, 1977. 



