159 



cotylédons are flesliy, sessile, linear-oblong obtuse. The 

 primary leaves are obovale-cuneaie tapering at the base 

 into a short wide stalk usually without any trace of stipules 

 but with a few stalked glands along the edges. The plu- 

 mule already shows a few pale yellow aculei. 



The mode of germination therefore does not differ 

 malerially from that of olher Roses in which I hâve 

 frequently (but by no means invariably), seen the first 

 one, two, or even ihree, leaves after the cotylédons 

 without stipules and with only a simple lamina. Higher 

 up the « adnate » stipules and a pair of leaflets are formed 

 in addition lo the terminal leaflet, still higher another 

 pair of leaflets appears and so on. 



Anatomy of the stipules and pétiole, — In an ordinary 

 Rose the stipules are, as it is called, « adnate », but 

 « enate » or « inseparaie » better expresses the real state 

 of the case. On culting across the pétiole and the stipules 

 emerging on either side from it, ihree vascular bundles 

 are seen, one central and one on either side thus : oQo. 

 Cords from the latéral bundles pass direcily into the sti- 

 pules, ail three of the main-bundies pass up the pétiole, 

 so that above, as well as helow the stipules there are three 

 cords, but at the very apex of the pétiole just beneath the 

 lower most pair of pinnae the cords form almost a closed 

 circle^, higher up the three cords again become separate 

 so that even the petiolule of the terminal leaflet i. e. the 

 end of the midrib shows 5 vascular bundles. 



In the pétiole of an ordinary Rose, beneath the epi- 

 dermis is a zone of cells containing chlorophyll, and 

 surrounding ihe ground tissue which is traversed by the 

 fibro-vascular cords before mentioned. In many Roses the 

 groove on the upper part of the pétiole is filled in with a 



