468 



Fishery Bulletin 93(3). 1 995 





EL. 

 <J 



e 

 a 



-Q 



INTERTIDAL 

 EELGRASS 



June Mid-June July 



SUBTIDAL 

 EELGRASS * 



August 



3 



a 



n 



to 



D 



o 



•a 

 c 

 w 



a 



fad 



ft. 

 U 

 e 



(S 





n 



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June Mid-June July August 



June Mid-June July August 



Month 



Figure 10 



Mean densities of staghorn sculpin, Leptocottus armatus, and Dungeness crab, Cancer magister (no./ha), from 

 intertidal and subtidal eelgrass and shell habitats, Grays Harbor, Washington, June-August 1989. Note dif- 

 ferent vertical scales. Each symbol represents the mean CPUE of 2 trawls. * = benthic habitat with shell, 

 ** = benthic habitat with wood and stick material. 



0+ Dungeness crab instars during a short period of 

 late spring and early summer settlement and growth 

 and that sculpin may be capable of exerting a pro- 

 nounced effect on crab year-class strength within 

 coastal estuaries (Armstrong, 1991). 



Acknowledgments 



We wish to acknowledge the Grays Harbor Sea Grant 

 Project for logistical and financial support. Thomas 

 Quinn, Donald Gunderson, Paul Dinnel, and Jose 

 Orensanz offered critical comments during all phases 

 of this work. Logistics and field sampling efforts were 

 contributed by Raul Palacios and Yun-Bing Shi. Greg 

 Williams contributed the use of unpublished data 

 pertaining to staghorn sculpin total to standard 

 length relationship and many engaging discussions 

 in the laboratory. Partial funding was provided by 

 Washington Sea Grant Program (Project No. R/SH-8 

 to David Armstrong). 



Literature cited 



Armstrong, J. L. 



1991. Food habits of staghorn sculpin, Leptocottus armatus, 

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 Cancer magister, in the Grays Harbor estuary, Washing- 

 ton. M.S. thesis, Univ. Washington, Seattle, WA, 88 p. 



Barshaw, D. E., and K. L. Lav alii. 



1988. Predation upon postlarval lobsters, Homarus 

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 eelgrass, mud and rocks. Mar. Ecol. Prog. Ser. 48:119-123. 



Bass, N. R., and A. E. Brafield. 



1972. The life cycle of the polychaete Nereis virens. Biol. 

 Assoc. (U.K.) 52:701-726. 

 Bayer, R. D. 



1985. Shiner perch and Pacific staghorn sculpins in Yaquina 

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1984. Fish diets and the benthic invertebrates in the estu- 

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 Can. Manuscript Rep. Fish. Aquat. Sci. No. 1799, 58 p. 

 Botsford, L., D. A. Armstrong, and J. Shenker. 



1989. Oceanographic influences on the dynamics of com- 

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